Bats (Chiroptera) in the collection of the Zoological Museum of Lviv University, Ukraine
The theriological collection of the Zoological Museum of Ivan Franko National University of Lviv was amassed during 142 years. It has over 3,800 exhibits, among which 343 (9 %) are bats (Chiroptera) that belong to 32 species. The bat collection is represented by study skins (254 exhibits) which are...
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nasplib_isofts_kiev_ua-123456789-1643912025-02-09T15:31:17Z Bats (Chiroptera) in the collection of the Zoological Museum of Lviv University, Ukraine Рукокрилі (Chiroptera) у колекції Зоологічного музею Львівського університету Shydlovskyy, I. Zatushevsky, A. Kusnezh, O. Фауна та систематика The theriological collection of the Zoological Museum of Ivan Franko National University of Lviv was amassed during 142 years. It has over 3,800 exhibits, among which 343 (9 %) are bats (Chiroptera) that belong to 32 species. The bat collection is represented by study skins (254 exhibits) which are being preserved in 22 special boxes. Stuffed and almost all fluid-preserved specimens of bats are presented only in the exposition. Besides, there are 6 skeletons and 11 skulls in the collection, which belong to at least 9 species of bats. The main part of the collection was gathered in Western Ukraine, and only a few samples originate from the South and East of Ukraine. In particular, the type series of Khazarian serotine Eptesicus lobatus Zagorodniuk, 2009 was brought from Eastern Ukraine. Tropical species are also presented in the collection: Hipposideros caffer, Epomophorus labiatus, Pteropus vampyrus, and species of the genera Neoromicia and Rousettus. The bat collection was amassed during 1900–2015. It widely represents the bat fauna of the Carpathian region and Transcarpathian lowland. Теріологічна колекція Зоологічного музею ЛНУ ім. І. Франка зібрана упродовж 142 років та налічує понад 3 800 музейних зразків, з яких 343 зразки, або 9 % — рукокрилі (Chiroptera) 32 видів. За типами музейних зразків колекція представлена тушками (254 екз.), які зберігаються у 22 спеціальних коробках. Опудала та майже всі мокрі препарати кажанів представлені в експозиції. Крім того, у колекції зберігаються шість скелетів та 19 черепів, що належать не менше ніж 9 видам кажанів. Основна частина колекції зібрана в межах західних областей України і лише поодинокими зразками представлено південь та схід. Зі cходу України, зокрема, є типовий матеріал пергача донецького, Eptesicus lobatus Zagorodniuk, 2009. Тропічні види представлені у колекції видами Hipposideros caffer, Epomophorus labiatus, Pteropus vampyrus, представниками родів Neoromicia та Rousettus. Хронологічно колекція зібрана упродовж 1900–2015 рр. Вона широко репрезентує хіроптерофауну регіону Карпат та Закарпатської низовини. 2018 Article Bats (Chiroptera) in the collection of the Zoological Museum of Lviv University, Ukraine / I. Shydlovskyy, A. Zatushevsky, O. Kusnezh // Theriologia Ukrainica. — 2018. — Т. 16. — С. 85–90. — Бібліогр.: 7 назв. — англ. 2616-7379 DOI: http://doi.org/10.15407/pts2018.16.085 https://nasplib.isofts.kiev.ua/handle/123456789/164391 599.4:069.5:5:378.4(477.83) en Theriologia Ukrainica application/pdf Національний науково-природничий музей НАН України |
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Фауна та систематика Фауна та систематика |
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Фауна та систематика Фауна та систематика Shydlovskyy, I. Zatushevsky, A. Kusnezh, O. Bats (Chiroptera) in the collection of the Zoological Museum of Lviv University, Ukraine Theriologia Ukrainica |
| description |
The theriological collection of the Zoological Museum of Ivan Franko National University of Lviv was amassed during 142 years. It has over 3,800 exhibits, among which 343 (9 %) are bats (Chiroptera) that belong to 32 species. The bat collection is represented by study skins (254 exhibits) which are being preserved in 22 special boxes. Stuffed and almost all fluid-preserved specimens of bats are presented only in the exposition. Besides, there are 6 skeletons and 11 skulls in the collection, which belong to at least 9 species of bats. The main part of the collection was gathered in Western Ukraine, and only a few samples originate from the South and East of Ukraine. In particular, the type series of Khazarian serotine Eptesicus lobatus Zagorodniuk, 2009 was brought from Eastern Ukraine. Tropical species are also presented in the collection: Hipposideros caffer, Epomophorus labiatus, Pteropus vampyrus, and species of the genera Neoromicia and Rousettus. The bat collection was amassed during 1900–2015. It widely represents the bat fauna of the Carpathian region and Transcarpathian lowland. |
| format |
Article |
| author |
Shydlovskyy, I. Zatushevsky, A. Kusnezh, O. |
| author_facet |
Shydlovskyy, I. Zatushevsky, A. Kusnezh, O. |
| author_sort |
Shydlovskyy, I. |
| title |
Bats (Chiroptera) in the collection of the Zoological Museum of Lviv University, Ukraine |
| title_short |
Bats (Chiroptera) in the collection of the Zoological Museum of Lviv University, Ukraine |
| title_full |
Bats (Chiroptera) in the collection of the Zoological Museum of Lviv University, Ukraine |
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Bats (Chiroptera) in the collection of the Zoological Museum of Lviv University, Ukraine |
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Bats (Chiroptera) in the collection of the Zoological Museum of Lviv University, Ukraine |
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bats (chiroptera) in the collection of the zoological museum of lviv university, ukraine |
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Національний науково-природничий музей НАН України |
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2018 |
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Фауна та систематика |
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https://nasplib.isofts.kiev.ua/handle/123456789/164391 |
| citation_txt |
Bats (Chiroptera) in the collection of the Zoological Museum of Lviv University, Ukraine / I. Shydlovskyy, A. Zatushevsky, O. Kusnezh // Theriologia Ukrainica. — 2018. — Т. 16. — С. 85–90. — Бібліогр.: 7 назв. — англ. |
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Theriologia Ukrainica, 16: 85–90 (2018)
http://doi.org/10.15407/pts2018.16.085
UDC 599.4:069.5:5:378.4(477.83)
BATS (CHIROPTERA) IN THE COLLECTION OF THE ZOOLOGICAL MUSEUM
OF LVIV UNIVERSITY, UKRAINE
I. Shydlovskyy1, A. Zatushevsky1, O. Kusnezh2
1 Zoological Museum, Ivan Franko National University of Lviv (Lviv, Ukraine)
2 Ukrainian Centre for Bat Protection, Lviv Division (Lviv, Ukraine)
Correspondence to: I. Shydlovskyy; Zoological Museum, Ivan Franko National University of Lviv, Hrushevsky St. 4,
Lviv, 79005 Ukraine;
e-mail: zoomus@franko.lviv.ua
Bats (Chiroptera) in the collection of the Zoological Museum of Lviv University, Ukraine. — I. Shyd-
lovskyy, A. Zatushevsky, O. Kusnezh. — The theriological collection of the Zoological Museum of Ivan
Franko National University of Lviv was amassed during 142 years. It has over 3,800 exhibits, among which
343 (9 %) are bats (Chiroptera) that belong to 32 species. The bat collection is represented by study skins
(254 exhibits) which are being preserved in 22 special boxes. Stuffed and almost all fluid-preserved specimens
of bats are presented only in the exposition. Besides, there are 6 skeletons and 11 skulls in the collection, which
belong to at least 9 species of bats. The main part of the collection was gathered in Western Ukraine, and only a
few samples originate from the South and East of Ukraine. In particular, the type series of Khazarian serotine
Eptesicus lobatus Zagorodniuk, 2009 was brought from Eastern Ukraine. Tropical species are also presented in
the collection: Hipposideros caffer, Epomophorus labiatus, Pteropus vampyrus, and species of the genera Neo-
romicia and Rousettus. The bat collection was amassed during 1900–2015. It widely represents the bat fauna of
the Carpathian region and Transcarpathian lowland.
Key words: bats, collections, zoological museum, Lviv University.
Рукокрилі (Chiroptera) у колекції Зоологічного музею Львівського університету. — І. В. Шидлов-
ський, А. Т. Затушевський, О. В. Кусьнеж. — Теріологічна колекція Зоологічного музею ЛНУ
ім. І. Франка зібрана упродовж 142 років та налічує понад 3 800 музейних зразків, з яких 343 зразки, або
9 % — рукокрилі (Chiroptera) 32 видів. За типами музейних зразків колекція представлена тушками
(254 екз.), які зберігаються у 22 спеціальних коробках. Опудала та майже всі мокрі препарати кажанів
представлені в експозиції. Крім того, у колекції зберігаються шість скелетів та 19 черепів, що належать
не менше ніж 9 видам кажанів. Основна частина колекції зібрана в межах західних областей України і
лише поодинокими зразками представлено південь та схід. Зі cходу України, зокрема, є типовий матері-
ал пергача донецького, Eptesicus lobatus Zagorodniuk, 2009. Тропічні види представлені у колекції вида-
ми Hipposideros caffer, Epomophorus labiatus, Pteropus vampyrus, представниками родів Neoromicia та
Rousettus. Хронологічно колекція зібрана упродовж 1900–2015 рр. Вона широко репрезентує хіроптеро-
фауну регіону Карпат та Закарпатської низовини.
Ключові слова: кажани, колекції, зоологічний музей, Львівський університет.
Introduction
The Zoological Museum of Ivan Franko National University of Lviv (ZMD) with its collections
is a unique asset of natural science, which makes possible to accumulate materials that cannot be
collected in a short period of time. It also ensures the preservation of collection samples of different
age, as well as animals collected in different parts of their range. At the same time, it allows process-
ing materials related to rare and endangered species, which populations are too small to be investi-
gated even by temporal extraction of individuals from the natural environment. Due to this feature,
museums (collections) provide the opportunity to fully study and compare the samples that cannot be
obtained or collected during a scientific course study or a research project, which are often limited
I. Shydlovskyy, A. Zatushevskyy, O. Kusnezh
86
by a small timeframe, poor funding, or the objects to reach are in different and often remote parts of
the range.
One of these animal groups are the chiropterans that carry out a hidden, nightfall or night activ-
ity. Moreover, the places of their reproduction are so concealed that special equipment is often
needed to finding them. In addition, all representatives of this group of the mammalian fauna of
Ukraine are listed in the Red Data Book of Ukraine (2009).
The purpose of this paper is to show the quantitative and qualitative changes in the collection of
bats that occurred during the 15-year period since the publication of its first synopsis (Bashta,
Shydlovskyy, 2001) and the characterization of the collection published during the study of rare rep-
resentatives of this order (Kusnezh, 2014). The electronic database of mammalian collections is be-
ing continuously updated for analysis (since 2010 in Excel format). It is a convenient tool for proc-
essing various data on individual museum objects and their series, as well as for searching and sum-
marizing of specific information.
The history of the collection’s content and volume
Reflecting on the history of mammalian species composition, it should be mentioned that the
theriological collection of ZMD has been enriched for 142 years and it includes over 3,800 museum
specimens, 343 or 9 % of which are bats (Chiroptera).
Analyzing the collection of bats, we can claim its gradual enrichment in both quality and quan-
tity. In particular, compared to 2001, when the revision of this collection was carried out, its abun-
dance increased from 125 study skins and mounted specimens of 14 species to 194 museum samples
of 25 species in 2010 (Zatushevskyy at al., 2010), and to 343 samples of 32 species by the end of
2016. This is without consideration of two unidentified specimens of Chiroptera, three unidentified
specimens of the genus Neoromicia, two unidentified juveniles of Pipistrellus sp., and one specimen
of Rousettus sp.
There are also 6 skeletons and 19 skulls of not less than 9 species in the bat collection, including
skeletons of such species as the common noctule (Nyctalus noctula, Schreber, 1774) — 2 specimens,
greater mouse-eared bat Myotis myotis (Borkhausen, 1797) — 1, pond bat M. dasycneme (Boie,
1825) — 1, Daubenton’s bat M. daubentonii (Kuhl, 1817) — 1, and an unidentified Pipistrellus sp.
Besides, there are skulls (1 specimen of each species) of the barbastelle Barbastella barbastellus
(Schreber, 1774), brown long-eared bat or common long-eared bat, Plecotus auritus (Linnaeus,
1758), Natterer’s bat Myotis nattereri (Kuhl, 1817), Nathusius’ pipistrelle Pipistrellus nathusii
(Keyserling and Blasius, 1839), as well as 3 specimens of the serotine bat Eptesicus serotinus
(Schreber, 1774), 5 specimens of Nyctalus noctula, and 7 specimens of Eptesicus lobatus (Zagorod-
niuk, 2009).
The collection is represented mainly by study skins that are stored in special cardboard boxes
sorted by species (Fig. 1 a). In particular, museum samples of this type are represented by 258 study
skins, among which 104 with skulls, in 22 boxes.
Mounted bats are only in the exhibition and they represent the bat fauna of the western regions
of Ukraine (common pipistrelle, Pipistrellus pipistrellus (Schreber, 1774), common bent-wing bat,
Miniopterus schreibersii (Kuhl, 1817), parti-colored bat, Vespertilio murinus Linnaeus, 1758, Geof-
froy’s bat, M. emarginatus (E. Geoffroy, 1806). There is only one specimen from Romania, namely
of the greater noctule bat, Nyctalus lasiopterus (Schreber, 1780), which is in the museum display.
Fluid-preserved specimens represent the fauna of tropical countries, in particular of Africa (Tri-
aenops persicus (Dobson, 1871), Hipposideros commersoni (E. Geoffroy, 1813), Sundevall’s round-
leaf bat, H. caffer (Sandevall, 1846), Epomophorus labiatus (Temminck, 1837), Neoromicia nanus
stampflii (Jentink, 1888) and Neoromicia sp.) and Southeastern Asia (the large flying fox, Pteropus
vampyrus (Linnaeus, 1758) from Sri Lanka). A specimen of the genus Rousettus, distributed mainly
in the southern hemisphere, is also exhibited.
In addition, a small amount of mummies and fluid-preserved specimens are also stored in the
stock collection (Table 1).
Bats (Chiroptera) in the collection of the Zoological Museum of Lviv University
87
Collection from Ukraine
The majority of specimens in the bat exhibition were collected in Ukraine. In particular, bats
from 7 of 24 oblasts and the Autonomous Republic of Crimea are represented. However, the quanti-
tative distribution of samples’ origin is not quite homogeneous. The largest collection came from
Lviv and Zakarpattia oblasts (42.3 % and 31.1 %, respectively), whereas such administrative units as
the Autonomous Republic of Crimea and Ivano-Frankivsk oblast are represented only by a single
specimen each. There are no samples from Chernivtsi and Khmelnytskyi oblasts. Concluding on the
collection’s record localities, the bulk of the collection is represented by specimens from the western
regions and only few specimens from the southern (e.g., AR Crimea) and eastern regions of Ukraine
(e.g., 8 item from Luhansk oblast). However, specimens from Luhansk oblast scientifically represent
the most valuable part of the collection, including type series of Eptesicus lobatus (Zagorodniuk,
2009; Eptesicus lobatus Zagorodniuk, 2010) with a holotype and five paratypes (table 2).
Chronology and geography of the collection
Chronologically, the museum’s collection of bats was amassed during 1902–2015. The very
first specimen of the collection, a greater noctule bat, is dated to December 1902 (Fig. 1 b). This is
the only specimen dated to that time. This very specimen has repeatedly drawn attention of local
scientists since Prof. K. Tatarinov (1956) wrote about it with the label “Opillya” stored in the collec-
tion of Lviv University. However, on the stand of the only copy available in our collection, on the
left is a black ink indication, which states the name of the species “Vesperugo vespexus noctula” and
its origin “18/12/1902 Dobrudza” and the composition “czaszka osobno” — the skull separately (in
Polish). To the right of this indication is a stock-keeping unit, the Latin name “N. siculus,” and the
signature of K. Tatarinov made with pencil, apparently later than the original inscription. Between
the Latin name and the professor’s signature there is an illegible Polish inscription with small and
narrow font, made by purple ink (“Polonensi?”). Therefore, the fact of discovery of the greater noc-
tule bat in the West of Ukraine remains obscure.
In the monograph “The Mammals of Western Ukraine” (Tatarinov, 1956), at the end of the es-
say the author writes on the greater noctule bat the following: “Apparently, this kind of noctule bat
in the western regions of the Ukrainian SSR is very rare, but in Dobrudja, according to verbal evi-
dence that we have, it is [a] common [species]". In our opinion, this is another confirmation that our
specimen of the greater noctule bat comes from the territory of Romania, namely from Dobrudja,
and therefore it was not discovered in western Ukraine.
a b
Fig. 2. Bats in the collection of ZMD: a — one of the oldest specimens of bats in the Museum, Nyctalus lasiopterus;
b — a box with study skins of bats as an example of their storing in the Museum. Photo by A. Zatushevsky.
Рис. 2. Кажани в колекції ЗМД: a — один із найстаріших зразків кажанів музею, Nyctalus lasiopterus; b —
коробка з тушками кажанів як приклад їхнього збереження у музеї. Фото А. Затушевського.
I. Shydlovskyy, A. Zatushevskyy, O. Kusnezh
88
Table 1. The number of different types of specimens in the museum’s bat collection.
Таблиця 1. Кількість різних типів зразків хіроптерологічної колекції музею
Type of museum specimen Amount Type of museum specimen Amount
Mounted specimens 14 Mummies 23
Study skins 258 Skeletons 6
Fluid-preserved specimens 23 Skulls 19
Table 2. Administrative-territorial representation of specimens of bats from Ukraine
Таблиця 2. Адміністративно-територіальна представленість зразків кажанів з території України
Admin. region Amount Admin. region Amount Admin. region Amount
Volyn 14 Luhansk 8 Ternopil 31
Zakarpattia 84 Lviv 113 AR Crimea 1
Ivano-Frankivsk 2 Rivne 16 No specified place* 8
* Including samples without label.
Table 3. Representativeness of the bat collection according to the geographical origin of specimens
Таблиця 3. Репрезентативність хіроптерологічної колекції за географічним походженням зразків
Place of origin Amount Place of origin Amount
Angola (Cabinda Province, Landana) 1 Romania (Dobruja) 1
Greece 1 Ukraine 292
Kenya 23 Sri-Lanka 1
Liberia 4 Africa (“Mapunga”) 1
Madagascar 1 no specified place* 16
Germany (Thüringen) 2 * Including samples without a label.
During 1909–1910, 19 mummies of the Sundevall’s leaf-nosed bat were received, brought by
Jerzy Wodzicki from the Great Rift Valley, Kenya. The next enrichment of the collection took place
only in 1935–1936, when Professor Jan Hirschler brought four specimens of Neoromicia sp. from
Liberia, three of which are still unidentified, and one identified as Neoromicia nanus stampflii. Later
on, the museum received a specimen of Nathusius’ pipistrelle (Pipistrellus nathusii) collected during
World War II by an unknown person in September 1942 in Greece.
The first collection of bats in the museum began to form in 1947 after World War II. However,
by 1960 specimens were received occasionally. T. D. Maznova first started systematically collect
Chiropterans in the caves of Ternopil oblast, in particular, near Korolivka, Bilche Zolote, and Uhryn
villages in the late 1960’s. After 1961, other collectors have joined the collection process and the bat
collection increased in amount to 88 specimens of 10 specimens within ten years. Although, after
1970, there was a significant suspension. The museum did not receive new samples of bats until the
late 20th century (namely until 1999).
In terms of geographical distribution, as it was already noted, the bat collection is represented,
mainly by Ukrainian samples. Only 35 specimens were collected in other countries (continents), in
particular 29 in Africa, 1 in Southeastern Asia, 3 in Western Europe, 1 in Central Europe, 1 in
Madagascar, and 292 in Eastern Europe (Table 3).
The new “era” in the bat collection’s enrichment dates back to the beginning of the 21st cen-
tury, when students began to conduct their coursework and graduation theses on bats under the guid-
ance of Associate Professors Eugenia Srebrodolska and Igor Dykyy. The activity in this area in-
creased and, as a result, dead bats found by students eventually were transferred to the museum’s
collection. Evidently, after 1999–2000 only two specimens arrived to the collection — a Brandt’s bat
(Myotis brandtii) from the southern suburbs of Lviv city and a common pipistrelle (Pipistrellus
pipistrellus) from Haivka village of Shatsk raion, Volyn oblast.
Bats (Chiroptera) in the collection of the Zoological Museum of Lviv University
89
During 2001–2015, after the first revision of the collection (Bashta, Shydlovskyy, 2001), its
volume increased significantly with 135 new specimens of 20 species.
Review on new samples
New samples obtained during 2001–2007 are characterized by a small amount — up to four-
five specimens per year. However, after 2008, the number of bat biomaterials that enriched the mu-
seum’s collection increased to an average of 13 specimens per year, with the highest number in
2014. The annual enrichment of the museum’s bat collection for the past 15 years is presented on
Fig. 2.
The largest number of study skins (47 items) belongs to specimens of the common noctule. One
of them was passed to the museum in 2002, and it was collected by V. Mysyuk at the Biological and
Geographical Research Station of Ivan Franko National University of Lviv, which is located within
Shatsk National Nature Park. Three other specimens were transferred from Uzhhorod in 2011 by
Y. Zizda, while the rest of specimens (43) was collected by I. Ivashkiv, O. Kusnezh, and M. Skyrpan
in Lviv during 2011–2014. These researchers discovered a place of mass death of bats that occured
at the beginning of their spring migration. This place was a hole on the roof of an old house entrance
through which the bats would depart. It was located above a pipe from which the funnel of rain and
melt-water usually fell after winter. That was the place where young bats used to fall into, while the
bottom of the pipe had still been frozen. Chiropterologists had repeatedly talked to the residents of
this house about changing the shape of the drainage funnel, which was made only three years after.
However, during this time more than 100 specimens of the common noctule died. Part of this
material is still being stored in the freezer chamber of the museum.
The second largest enrichment was by specimens of the serotine bat (Eptesicus serotinus). Dur-
ing the period of studies, 28 specimens were obtained, almost half of which (13) was collected in
2010 by E. Stetskiv and I. Ivashkiv. One or two other specimens of this species, collected in Rivne
and Lviv oblasts, were added to the collection as well.
In 2009, the museum’s chiropterological collection was enriched with barbastelle specimens
(five mummies) prepared by A.-T. Bashta and I. Ivashkiv and collected in the outskirts of Tarakaniv
village, Dubensky district, Rivne oblast.
In 2012, one of the authors of this article (O. Kusnezh) enriched the collection of bats with four
specimens of Bechstein’s bat (Myotis bechsteini), three of which were collected in Ternopil oblast
(Kalaharivka village, Gusyatyn district) and one in the Roztochya Biosphere Reserve (Ivano-
Frankove, Yavorivsky district, Lviv oblast).
Valuable samples
Thus, the bat collection of ZMD widely represents the bat fauna of the Ukrainian Carpathians
and Transcarpathian lowland. Among valuable samples of the collection, it is worth listing the mate-
rials from the exposition and stocks that evidence the presence of certain species of bats in Western
Ukraine in the middle of the 20th century.
0
10
20
30
2001 2003 2005 2007 2009 2011 2013 2015
number of specimens
Fig. 2. The number of specimens received by
the museum and included into the bat collec-
tion during 2001–2015.
Рис. 2. Кількісне представлення надхо-
джень кажанів до хіроптерологічної коле-
кції музею протягом 2001–2015 рр.
I. Shydlovskyy, A. Zatushevskyy, O. Kusnezh
90
Among them are the whiskered bat (Myotis mystacinus), the lesser mouse-eared bat (Myotis
blythii), and the lesser horseshoe bat (Rhinolophus hipposideros) collected in 1947–1960, as well as
the common bent-wing bat (Miniopterus schreibersii) (47 specimens) and a number of other species
collected in 1961–1962 in Zakarpattia oblast.
Scientifically valuable is the collection of type series of Eptesicus lobatus, which may serve for
future examinations of the genus Eptesicus, which is widespread in Eastern Europe, as well as for
morphological studies of the post-calcarial lobe of bats and other bat fauna research of the Azov and
Donetsk Uplands too.
Equally interesting are collection specimens that represent the tropical fauna and are real evi-
dence of certain species’ presence within their range. These findings also provoke new questions for
the researchers. In particular, regarding to representatives of the genus Neoromicia, one of which,
according to our considerations, belongs to the species N. nanus stampflii. However, on the label of
our specimen the species is listed as N. stampflii, i.e. the species is identified in the “Mammal Spe-
cies of the World” (Wilson, Reeder, 2005) as extinct. Nevertheless, our specimens may belong to
other three species or subspecies.
References • Література
Bashta, А.-Т., I. Shydlovsky. 2001. Collection of bats (Chirop-
tera) in B. Dybowski Zoological Museum of the Lviv Na-
tional University. Scientific Notes of State Natural History
Museum. Lviv, 16: 41–45. (In Ukrainian)
[Башта, А.-Т. В., І. В. Шидловський. 2001. Колекція ру-
кокрилих (Chiroptera) Зоологічного музею ім. Б. Дибов-
ського Львівського національного університету. Наукові
записки Державного природознавчого музею. Львів, 16:
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Donetsk... 2010. Donetsk Serotine (Eptesicus lobatus Zagorod-
niuk, 2009) — a new species of bats. Stanislavivskyy Natu-
ralist (web-site). https://goo.gl/yECJvR (In Ukrainian)
[Пергач... 2010. Пергач донецький (Eptesicus lobatus Za-
gorodniuk, 2009) — новоописаний вид кажанів. Станіс-
лавівський натураліст (веб-сайт). https://goo.gl/yECJvR]
Kusnezh, A. V. 2014. Rare species of bats (Chiroptera) in the
collections of the Zoological Museum of Ivan Franko Na-
tional University of Lviv. In: Zagorodniuk, I. (ed.). Zoologi-
cal Collections and Museums. National Museum of Natural
History, Ukr. Acad. Sci. Kyiv, 63–66. (In Ukrainian)
[Кусьнеж, О. В. 2014. Рідкісні види рукокрилих (Chirop-
tera) у фондах Зоологічного музею Львівського НУ ім.
І. Франка. Загороднюк, І. (ред.). Зоологічні колекції та
музеї: Зб. наук. пр. Національний науково-природничий
музей НАН України. Київ, 63–66.]
Tatarynov, K. A. 1956. Mammals of the Western Regions of
Ukraine. Vydavn. AS Ukr. SSR, Kyiv, 1–186. (In Ukrainian)
[Татаринов, К. А. 1956. Звірі західних областей України.
Вид-во АН УРСР, Київ, 1–186.]
Zagorodniuk, I. 2009. Morphology of post-calcarial lobe in bats
and its variation in Eptesicus “serotinus” (Mammalia). Vis-
nyk of the Lviv University. Series Biology, 51: 157–175. (In
Ukrainian)
[Загороднюк, І. 2009. Морфологія епіблеми у кажанів та
її мінливість у Eptesicus «serotinus» (Mammalia). Вісник
Львівського національного університету. Серія Біоло-
гічна, 51: 157–175.]
Zatushevskyy, A. T., I. V. Shydlovskyy, O. S. Zakala, I. V.
Dykyy, O. V. Holovachov, M. A. Senyk, Kh. J. Romanova.
2010. Catalogue of the Mammals Collection of the Zoologi-
cal Museum of Ivan Franko National University of Lviv.
Publishing Center of the Ivan Franko National University of
Lviv, Lviv, 1–442. (In Ukrainian)
[Затушевський, А. Т., І. В. Шидловський, О. С. Закала,
І. В. Дикий, О. В. Головачов, М. А. Сеник, Х. Й. Рома-
нова. 2010. Каталог колекцій ссавців Зоологічного музею
Львівського національного університету імені Івана
Франка. Вид. центр ЛНУ ім. І. Франка, Львів, 1–442.]
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/ENU (Use these settings to create Adobe PDF documents best suited for high-quality prepress printing. Created PDF documents can be opened with Acrobat and Adobe Reader 5.0 and later.)
>>
/Namespace [
(Adobe)
(Common)
(1.0)
]
/OtherNamespaces [
<<
/AsReaderSpreads false
/CropImagesToFrames true
/ErrorControl /WarnAndContinue
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/IncludeNonPrinting false
/IncludeSlug false
/Namespace [
(Adobe)
(InDesign)
(4.0)
]
/OmitPlacedBitmaps false
/OmitPlacedEPS false
/OmitPlacedPDF false
/SimulateOverprint /Legacy
>>
<<
/AddBleedMarks false
/AddColorBars false
/AddCropMarks false
/AddPageInfo false
/AddRegMarks false
/ConvertColors /ConvertToCMYK
/DestinationProfileName ()
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/Downsample16BitImages true
/FlattenerPreset <<
/PresetSelector /MediumResolution
>>
/FormElements false
/GenerateStructure false
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/MultimediaHandling /UseObjectSettings
/Namespace [
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(CreativeSuite)
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]
/PDFXOutputIntentProfileSelector /DocumentCMYK
/PreserveEditing true
/UntaggedCMYKHandling /LeaveUntagged
/UntaggedRGBHandling /UseDocumentProfile
/UseDocumentBleed false
>>
]
>> setdistillerparams
<<
/HWResolution [2400 2400]
/PageSize [612.000 792.000]
>> setpagedevice
|