New Species of the Genus Lasioderma (Coleoptera, Anobiidae) from Crimea and the Caucasus
Five species of the genus Lasioderma Stephens, 1835 from Crimea (L. dolini sp. n., L. tauricum sp. n.), North Caucasus (L. prolixum sp. n., L. sparsum sp. n.), and Armenia (L. morulum sp. n.) are described. A key to 17 species of the genus Lasioderma occurring in these regions is compiled. Описаны 5...
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| Дата: | 2011 |
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Інститут зоології ім. І.І. Шмальгаузена НАН України
2011
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| Цитувати: | New Species of the Genus Lasioderma (Coleoptera, Anobiidae) from Crimea and the Caucasus / I.N. Toskina // Вестник зоологии. — 2011. — Т. 45, № 3. — С. 195–207. — Бібліогр.: 13 назв. — англ. |
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Digital Library of Periodicals of National Academy of Sciences of Ukraine| _version_ | 1860178812514336768 |
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| author | Toskina, I.N. |
| author_facet | Toskina, I.N. |
| citation_txt | New Species of the Genus Lasioderma (Coleoptera, Anobiidae) from Crimea and the Caucasus / I.N. Toskina // Вестник зоологии. — 2011. — Т. 45, № 3. — С. 195–207. — Бібліогр.: 13 назв. — англ. |
| collection | DSpace DC |
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| description | Five species of the genus Lasioderma Stephens, 1835 from Crimea (L. dolini sp. n., L. tauricum sp. n.), North Caucasus (L. prolixum sp. n., L. sparsum sp. n.), and Armenia (L. morulum sp. n.) are described. A key to 17 species of the genus Lasioderma occurring in these regions is compiled.
Описаны 5 видов рода Lasioderma Stephens, 1835 из Крыма (L. dolini sp. n., L. tauricum sp. n.), Северного Кавказа (L. prolixum sp. n., L. sparsum sp. n.) и Армении (L. morulum sp. n.). Составлена определительная таблица 17 видов рода Lasioderma для названного региона.
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| first_indexed | 2025-12-07T18:01:06Z |
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| fulltext |
UDC 395.766.4(47)
NEW SPECIES OF THE GENUS LASIODERMA (COLEOPTERA,
ANOBIIDAE) FROM CRIMEA AND THE CAUCASUS
I. N. Toskina
107241, post box 25, Moscow, Russia
E-mail: nina_11235813@mail.ru
Received 12 October 2010
Accepted 30 March 2011
New Species of the Genus Lasioderma (Coleoptera, Anobiidae) from Crimea and the Caucasus. Toski -
na I. N. – Five species of the genus Lasioderma Stephens, 1835 from Crimea (L. dolini sp. n., L. tau-
ricum sp. n.), North Caucasus (L. prolixum sp. n., L. sparsum sp. n.), and Armenia (L. morulum sp. n.)
are described. A key to 17 species of the genus Lasioderma occurring in these regions is compiled.
Ke y wo r d s: Coleoptera, Anobiidae, Xyletininae, Lasioderma, Crimea, Caucasus, new species, key.
Íîâûå âèäû ðîäà Lasioderma (Coleoptera, Anobiidae) èç Êðûìà è Êàâêàçà. Òîñêèíà È. Í. –
Îïèñàíû 5 âèäîâ ðîäà Lasioderma Stephens, 1835 èç Êðûìà (L. dolini sp. n., L. tauricum sp. n.),
Ñåâåðíîãî Êàâêàçà (L. prolixum sp. n., L. sparsum sp. n.) è Àðìåíèè (L. morulum sp. n.).
Ñîñòàâëåíà îïðåäåëèòåëüíàÿ òàáëèöà 17 âèäîâ ðîäà Lasioderma äëÿ íàçâàííîãî ðåãèîíà.
Êëþ÷åâûå ñ ëîâ à: Coleoptera, Anobiidae, Xyletininae, Lasioderma, Êðûì, Êàâêàç, íîâûå âèäû,
òàáëèöà äëÿ îïðåäåëåíèÿ âèäîâ.
Introduction
Anobiid beetles of the genus Lasioderma were studied in Crimea and the Caucasus by A. Reitter (1884,
1913), who described L. punctulatum Reitter, 1884 and L. babadjanidesi Reitter, 1913. Approximately at the
same time, Schilsky (1899) described L. kiesenwetteri Schilsky, 1899, and Pic (1904) gave very short descrip-
tion of L. redtenbacheri var. caucasicum Pic, 1904. Later, F. Español (1970, 1992) figured and commented
L. haemorrhoidale (Illiger, 1807) and S. M. Iablokoff-Khnzorian (1976) illustrated and gave new records of
L. aterrimum Roubal, 1916. V. D. Logvinovskij (1977) described L. kryzhanovskii Logvinovskij, 1977 and pro-
vided a key to Lasioderma species of the USSR in his “Fauna of USSR” Anobiidae monograph (Logvinovskij,
1985). Recently, I. N. Toskina described L. mazokhini Toskina, 1999, L. mikheechevi Toskina, 1999 and
L. oculeum Toskina, 1999 and provided the most comprehensive key to the species of this genus occurring in
the USSR and adjacent countries. In this work, 5 additional new species from Crimea and the Caucasus of
the genus Lasioderma are described, and a key to species occurring in Crimea and Caucasus regions is pro-
vided.
Material and methods
Length of the pronotum was measured in profile in a position with the apical and basal margins aligned
at the same level. Measurements of the pronotum from above give higher error. The elytral length was mea-
sured from the basal margin of scutellum along elytral suture. The elytral width was measured somewhat pos-
teriorly of the shoulders. Aedeagus is figured from its dorsal side, and the pseudopositors from their ventral
side. In cases when hind tarsi are lost the ratio of the segments in the middle or fore tarsi is used, and then
the ratio of tibia and tarsus lengths is not used. Measurements with asterisk (*) are those of the holotype.
Material is deposited in the collections of Hungarian Museum of Natural History (Budapest)
(HMNH), the Zoological Museum of Moscow University (ZMUM), and Zoological Institute of the Russian
Academy of Sciences, St.-Petersburg (ZISP).
Vestnik zoologii, 45(3): 195—207, 2011 Ôàóíà è ñèñòåìàòèêà
Descriptions of new species
Lasioderma dolini sp. n. (fig. 1, 1—10)
Ma t e r i a l. Holotype }: USSR, [Crimea], Kizil-Dzhar, 8.05.1975 (Dolin) (HMNH). 2 paratypes with
the same label data (HMNH, ZMUM).
De s c r i p t i o n. Habitus as on fig. 1,1. Dark yellow beetles; antenna light yellow.
Pubescence light yellow, homogenous, fine, appressed on elytral disc, raised on sides.
196
Fig. 1. Lasioderma dolini sp. n., holotype }: 1 – general outline of body; 2 – antenna; 3 – maxillary pal-
pus; 4 – anterior part of body, lateral view; 5 – pronotum, dorsal view (with pubescence pattern); 6 –
punctation on pronotal disc; 7 – structure of elytral surface; 8 – the middle of metasternum and 1st visible
abdominal sternite; 9 – fore tarsus (part); 10 – pseudopositor: coxite top with style. Scale bar: 1.0 mm (1);
0.5 mm (4, 5, 8); 0.2 mm (2, 9); 0.1 mm (3, 6, 7); 0.05 mm (10).
Ðèñ. 1. Lasioderma dolini sp. n., ãîëîòèï }: 1 – îáùèé êîíòóð òåëà; 2 – óñèê; 3 – ïîñëåäíèé ÷ëåíèê
÷åëþñòíûõ ùóïèêîâ; 4 – ïåðåäíÿÿ ÷àñòü òåëà, âèä ñáîêó; 5 – ïåðåäíåñïèíêà, âèä ñâåðõó (ñ ðèñóí-
êîì îïóøåíèÿ); 6 – ïóíêòèðîâêà íà äèñêå ïåðåäíåñïèíêè; 7 – ñòðóêòóðà ïîâåðõíîñòè íàäêðûëüåâ;
8 – ñåðåäèíà çàäíåãðóäè è 1-ãî âèäèìîãî áðþøíîãî ñòåðíèòà; 9 – ïåðåäíÿÿ ëàïêà (÷àñòü); 10 –
ëîæíûé ÿéöåêëàä: âåðøèíà êîêñèòà ñî ñòèëåì. Ìàñøòàáíûå ëèíåéêè: 1,0 ìì (1); 0,5 ìì (4, 5, 8);
0,2 ìì (2, 9); 0,1 ìì (3, 6, 7); 0,05 ìì (10).
Head. Frons slightly convex. Eyes round, convex, separated by 1.8 eye diameters.
Antenna: 3rd segment almost as long as the 2nd one; 4th segment with oblique and
concave upper margin; 5—10th segments with concave upper margin and convex late -
ral one. 11th segment 3.3 times as long as wide and 1.6 times as long as the 10th seg-
ment. All segments oblong (fig. 1, 2). Last segment of maxillary palpus is spindle-
shaped, 3.3 times as long as thick, with oblique truncate top and with carina on it
(fig. 1, 3).
P r ono t um 1.5—1.6* times as wide as long, with base arcuate, anterior angles
rectangular, posterior ones not developed (fig. 1, 4). Pronotum slightly convex, lateral
margins narrowly flattened, sides very slightly bulging. Pubescence not parted to two
sides (fig. 1, 5). Surface with dual punctation, small punctures very fine (fig. 1, 6).
S c u t e l l um triangular. Elytra 1.8*—2.0 times as long as wide and 2.8*—3.0 times
as long as pronotum. Surface very gently wrinkled (fig. 1,7).
Me t a s t e r num nearly as long as 1st visible abdominal sternite. Anterior rims a
little approximated and evenly rounded medially. Medial portion significantly stronger
convex in distal third of metasternum (fig. 1, 8). Surface finely wrinkled with puncture
traces.
L e g s. Fore tibiae not widening to their apices. 1st segment of fore tarsus 1,7 times
as long as 2nd; 2nd segment 1.7 times as long as 3rd (fig. 1, 9). Other tarsi lost or
strongly damaged in type specimens.
P s e udopo s i t o r. Style looking as elongated cone with long chaeta on its top;
coxite top with rather long chaetae (fig. 1, 10).
Length 4.5*—5.0 mm, width 1.8—1.9* mm.
D i f f e r e n t i a l d i a g no s i s. L. dolini differs from other species of the genus
Lasioderma with having yellow-rufous or reddish-brown colour and with pubescens not
parted to two sides on pronotum occurring in this region (L. haemorrhoidale,
L. kryzhanovskii, L. mazokhini, L. oculeum, L. prolixum sp. n., L. punctulatum and
L. serricorne (Fabricius, 1792) as follows. L. dolini differs from L. serricorne and L. pro-
lixum by the non-dilated fore tibiae (fore tibiae dilated to their apices in the compared
species); it differs from L. haemorrhoidale, L. mazokhini, and L. oculeum by evenly
rounded anterior rims of metasternum (rims approximated in the middle and straigh-
tened or curved forwards in the compared species); the new species differs from
L. kryzhanovskii by homogenous pubescence on elytra (in the compared species,
pubescence dual on elytra) (see Logvinovskij, 1985: 134). At last, L. dolini differs from
L. punctulatum in fine punctation (in L. punctulatum, punctation is already visible with
magnification x10).
E t ymo l o g y. The new species is named in memory of Ukrainian entomologist
Vladimir Gdalich Dolin (1932—2004), who collected the type series.
Lasioderma morulum sp. n. (fig. 2, 1—10)
Ma t e r i a l. Holotype {: “USSR, Armenia, Gokh-Guekhard, 1500 m, 30 km E from Yerevan,
29.05.1980, N 129” (J. Papp) (HMNH).
De s c r i p t i o n. Habitus as on fig. 2, 1. Black beetle except dark yellow elytral
apices; continued as narrow stripe along lateral margin of elytron to metasternum.
Pubescence very fine, dark, homogenous, appressed on elytra.
He ad. Frons almost flat. Eyes round, moderately convex, separated by 2.7 eye
diameters. Antenna: 3rd segment 1.5 times as long as the 2nd one; 4—5th segments with
oblique and concave upper margin and convex lateral one; 6—9th segments with straight
upper and lateral margins (10—11th segments are lost). All segments except 5th oblong
(fig. 2, 2).
197
P rono t um 1.5 times as wide as long; anterior angles rectangular, posterior angles
slightly developed (fig. 2, 3). Sides not bulging, lateral margins not flattened.
Pubescence not parted to two sides (fig. 2, 4). Surface with dual punctation, looking as
if pitted because of large punctures (fig. 2, 5).
S c u t e l l um triangular. Elytra 1.5 times as long as wide and 2.1 times as long as
pronotum. Surface with dual punctation and wrinkles (fig. 2, 6).
Me t a s t e r num 1.5 times as long as 1st visible abdominal sternite. Anterior rims
approximated and straightened medially. (fig. 2, 7).
L e g s. Fore tibiae not widened to their apices. Hind tarsus more than 0.8 times as
long as hind tibia; 1st segment 1.3 times as long as 2nd segment; 2nd segment 1.9 times
198
Fig. 2. Lasioderma morulum sp. n., holotype {: 1 – general outline of body; 2 – antenna; 3 – anterior part
of body, lateral view; 4 – pronotum, dorsal view (with pubescence pattern); 5 – punctation on pronotal
disc; 6 – punctation on elytral disc; 7 – the middle of metasternum and 1st visible abdominal sternite; 8 –
hind tibia and tarsus; 9 – aedeagus; 10 – distal part of genital ring. Scale bar: 1.0 mm (1); 0.5 mm (3, 4);
0.2 mm (2, 7—10); 0.1 mm (5, 6).
Ðèñ. 2. Lasioderma morulum sp. n., ãîëîòèï {: 1 – îáùèé êîíòóð òåëà; 2 – óñèê; 3 – ïåðåäíÿÿ ÷àñòü
òåëà, âèä ñáîêó; 4 – ïåðåäíåñïèíêà, âèä ñâåðõó (ñ ðèñóíêîì îïóøåíèÿ); 5 – ïóíêòèðîâêà íà äèñêå
ïåðåäíåñïèíêè; 6 – ïóíêòèðîâêà íà äèñêå íàäêðûëüåâ; 7 – ñåðåäèíà çàäíåãðóäè è 1-ãî âèäèìîãî
áðþøíîãî ñòåðíèòà; 8 – ãîëåíü è ëàïêà çàäíåé íîãè; 9 – ýäåàãóñ; 10 – äèñòàëüíûé êîíåö ãåíèòàëü-
íîãî êîëüöà. Ìàñøòàáíûå ëèíåéêè: 1,0 ìì (1); 0,5 ìì (3, 4); 0,2 ìì (2, 7—10); 0,1 ìì (5, 6).
as long as 3rd one; 3rd segment 1.3 times as long as 4th segment; 5th segment almost
as long as the 2nd one; claws large (fig. 2, 8).
A ed e a gu s. Penis narrows to its apex, 9.3 times as long as wide at apex; sudden-
ly widened in basal third. Internal penial sac with longitudinal row of 6—7 small spines
and with 3 spines of medium sizes; one of them probably representing end of short
bifurcated “brush”. Parameres shortly bifurcated at ends, outgrowths nearly reaching
paramere apices (fig. 2, 9). Cross-piece of distal end of genital ring looking like thin
band slightly widened at every side (fig. 2, 10).
Length 2.25 mm, width 1.10 mm.
D i f f e r e n t i a l d i a g no s i s. The new species differs from other of black-coloured
Lasioderma species occurring in examined region (L. aterrimum; L. obscurum (Solsky,
1867)) in the following characters. L. morulum differs from L. aterrimum in oblong
6—9th antennal segments (4—10th antennal segments strongly transverse in L. aterri-
mum (Iablokoff-Khnzorian, 1976, fig. 10)). L. morulum differs from L. obscurum in
round eyes and pubescence not parted to two sides on pronotum (eyes shortly oval and
pubescence parted to two sides on pronotum in L. obscurum).
E t ymo l o g y. Species name comes from the Latin morulus (black, dark, gloomy),
reflecting colour of its body.
Lasioderma prolixum sp. n. (fig. 3, 1—12)
Ma t e r i a l. Holotype {: [Russia,] Krasnodar Terr., Severskij Distr., Ubinskoe Forestry, 21.06.1971
(Nikitsky) (as “Lasioderma obscurum Sols.” M. Dubrovina det.) (ZMUM, coll. Nikitsky).
De s c r i p t i o n. Habitus as on fig. 3, 1. Beetle blackish-brown, tinged with dark
red; margins and elytral apices not light-coloured. Pubescence dark grey, fine, homoge-
nous, appressed.
He ad. Frons slightly convex. Eyes round, slightly convex, separated by about
2.5 eye diameters. Antenna: 1st segment unusually flattened; 3rd segment 1.4 times as
long as the 2nd one; 4—5th segments with oblique and concave upper margin and con-
vex lateral one; 6—9th segments strongly transverse, their upper and lower (i. e. later-
al) margins straight. (10th and 11th segment lost) (fig. 3, 2).
P r ono t um 1.47 times as wide as long; anterior angles slightly acute, posterior
ones very clear, not rounded (fig. 3, 3). Lateral margins not flattened. Pubescence not
parted to two sides (fig. 3, 4). Surface with dual punctation, and more large punctures
are nearly as large as small ones (fig. 3, 5). Difference between large and small punc-
tures is more distinct near basal margin (fig. 3, 6).
S c u t e l l um triangular with blunt top. Elytra 1.4 times as long as wide and
2.3 times as long as pronotum. Surface with fine, uniform punctation (fig. 3,7).
Me t a s t e r num 1.5 times as long as the 1st visible abdominal sternite; anterior
rims approximated in the middle, 1st rim is straight, 2nd one a little curved forwards
(fig. 3, 8).
L e g s. Fore tibiae dilated to their apices (fig. 3, 9); all tibiae flattened. Tarsi short:
hind tarsus 0.54 times as long as its tibia; 1st segment of middle tarsus 3 times as long
as the 2nd one; 2nd segment 1.2 times as long as the 3rd one; the latter 1.2 times as
long as the 4th one; 5th segment 1.4 times as long as the 2nd one (fig. 3, 10).
A ed e a gu s. Penis narrows to rounded apex, 7.2 times as long as wide before
rounded apex. Internal penial sac has apical and basal “brushes”; basal ends are com-
pleted with 3 small needle-shaped sclerites in apical “brush”. The apical end of basal
“brush” is completed with rather small digitate sclerite. Basal parts of parameres (from
dorsal surface) with separated lobes. Side outgrowth consists of two halves and reach-
es the paramere apex. Phallobase short (fig. 3, 11). Cross-piece of genital ring is rib-
bon-like, strongly curved (fig. 3, 12).
199
Length 3.20 mm, width 1.75 mm.
D i f f e r e n t i a l d i a g no s i s. The new species belongs to the group of yellow- or
red-brown beetles of the genus Lasioderma with pubescence not parted to two sides on
pronotum, fore tibiae dilated to their apices. L. prolixum differs from the cosmopolitan
200
Fig. 3. Lasioderma prolixum sp. n., holotype {: 1 – general outline of body; 2 – antenna; 3 – anterior part
of body, lateral view; 4 – pronotum, dorsal view (with pubescence pattern); 5 – punctation on pronotal
disc; 6 – punctation near pronotal basal margin; 7 – punctation on elytral disc; 8 – the middle of metaster-
num and 1st visible abdominal sternite; 9 – fore tibia; 10 – middle tarsus; 11 – aedeagus; 12 – distal part
of genital ring. Scale bar: 1.0 mm (1); 0.5 mm (3, 4, 8, 9); 0.2 mm (2, 10—12); 0.1 mm (5—7).
Ðèñ. 3. Lasioderma prolixum sp. n., ãîëîòèï {: 1 – îáùèé êîíòóð òåëà; 2 – óñèê; 3 – ïåðåäíÿÿ ÷àñòü
òåëà, âèä ñáîêó; 4 – ïåðåäíåñïèíêà, âèä ñâåðõó (ñ ðèñóíêîì îïóøåíèÿ); 5 – ïóíêòèðîâêà íà äèñêå
ïåðåäíåñïèíêè; 6 – ïóíêòèðîâêà ïåðåäíåñïèíêè ó áàçàëüíîãî êðàÿ; 7 – ïóíêòèðîâêà íà äèñêå íàä-
êðûëüåâ; 8 – ñåðåäèíà çàäíåãðóäè è 1-ãî âèäèìîãî áðþøíîãî ñòåðíèòà; 9 – ïåðåäíÿÿ ãîëåíü; 10 –
ñðåäíÿÿ ëàïêà; 11 – ýäåàãóñ; 12 – äèñòàëüíûé êîíåö ãåíèòàëüíîãî êîëüöà. Ìàñøòàáíûå ëèíåéêè:
1,0 ìì (1); 0,5 ìì (3, 4, 8, 9); 0,2 ìì (2, 10—12); 0,1 ìì (5—7).
L. serricorne by clearly developed posterior angles of the pronotum, in strongly trans-
verse middle antennal segments with straight lower margins, approximated anterior rims
of the metasternum, by very dark (nearly black) body colour (in L. serricorne, prono-
tum posterior angles not developed, middle antennal segments with convex lateral mar-
gin, anterior rims evenly rounded in metasternum, body greyish-red, antenna yellow).
These species differ also in penial armaments (see article by Español, 1972, fig. 33; and
fig. 3, 11 in this article).
E t ymo l o g y. Species name comes from the Latin prolixus (wide, broad, exten-
sive), reflecting its body shape.
Lasioderma sparsum sp. n. (fig. 4, 1—10)
Ma t e r i a l. Holotype {: [Russia, Krasnodar Terr.] Anapa, 27.07.1962 (V. Meshcheryakov) (ZMUM).
De s c r i p t i o n. Habitus as on fig. 4, 1. Beetle brown: head, pronotum and
abdomen black-brown, elytra reddish-brown, elytral apices with yellow-red spot.
Pubescence light grey, heterogeneous, raised, rather long, arranged with uneven and
unclear transverse stripes and spots on elytra. Elytral surface with 2—3 costae near
suture and lateral margins (fig. 4, 2).
He ad. Frons almost flat. Eyes round, convex, separated by 1.6 eye diameters.
Antenna: 3rd segment 1.5 times as long as 2nd; 4th segment with oblique and straight
upper and straight lateral margins; 5—10th segments with slightly convex lateral and
straight upper margin. 11th segment 3 times as long as wide and 1.5 times as long as
10th segment. All segments oblong (fig. 4, 3). Last segment of maxillary palpus spin-
dle-shaped.
P r ono t um 1.55 times as wide as long; anterior angles rectangular, posterior ones
slightly developed (fig. 4, 4). Pronotum flattened at posterior angles. Pubescence part-
ed to two sides and arranged unevenly on pronotum surface (fig. 4, 5). The latter with
uniform punctation, its punctures separated by 0.25—0.5 puncture diameters (fig. 4, 6).
S c u t e l l um semioval. Elytra 1.6 times as long as wide and 2.6 times as long as
pronotum. Surface with dual punctation, small punctures separated by 1—2 puncture
diameters (fig. 4, 7).
Me t a s t e r num twice as long as the 1st visible abdominal sternite; anterior rims
approximated a little in the middle and widely straightened (fig. 4, 8).
L e g s. Fore tibiae not dilated to the apices. 1st segment of middle tarsus 1.5 times
as long as the 2nd one; 2nd segment 1.8 times as long as the 3rd one; 3rd segment
1.1 times as long as the 4th one; 5th segment almost as long as the 2nd one (fig. 4, 9).
A ed e a gu s. Internal penial sac turned out, its armaments consists of longitudinal
row of 8 small spines, 2 detached small spines, and 3 large spines (fig. 4, 10).
Length 3.5 mm, width 1.65 mm.
D i f f e r e n t i a l d i a g no s i s. The new species differs from other species of the
genus Lasioderma without black body colouration, with pubescence parted to two sides
on pronotum, and distributed in the named region (L. babadjanidesi, L. redtenbacheri
(Bach, 1852), L. mikheechevi, L. kiesenwetteri, L. tauricum sp. n.) in the following char-
acters. L. sparsum has rims of metasternum which are approximated and straightened
in the middle (these rims evenly rounded in L. babadjanidesi, L. redtenbacheri; and rims
slightly approximated and 1st rim hardly straightened in L. mikheechevi). L. sparsum
differs from L. kiesenwetteri by brown colour of body and pubescence parted to two
sides on pronotal disc (body yellow-red and pubescence parted near apical margin of
pronotum in L. kiesenwetteri). L. sparsum differs from L. tauricum in tarsi structures
(2nd segment about as long as the 1st one, and 5th segment 1.7 times as short as the
2nd one in L. tauricum, whereas 1st segment 1.5 times as long as the 2nd one and 5th
segment as long as the 2nd one in L. sparsum). Also L. sparsum differs from the named
201
species in spotted pubescence on elytra and armaments of internal penial sac (see
Toskina, 1999: figs 44, 92, 140, Logvinovskij, 1985: fig. 228, and this paper: fig. 4,10
and 5,10).
202
Fig. 4. Lasioderma sparsum sp. n., holotype {: 1 – general outline of body; 2 – right elytron with pubescence
pattern; 3 – antenna; 4 – anterior part of body, lateral view; 5 – pronotum, dorsal view (with pubescence
pattern); 6 – punctation on pronotal disc; 7 – punctation on elytral disc; 8 – the middle of metasternum
and 1st visible abdominal sternite; 9 – middle tarsus; 10 – internal penial sac (turned out). Scale bar:
1.0 mm (1); 0.5 mm (2, 4, 5, 8); 0.2 mm (3, 9); 0.1 mm (6, 7, 10).
Ðèñ. 4. Lasioderma sparsum sp. n., ãîëîòèï {: 1 – îáùèé êîíòóð òåëà; 2 – ïðàâîå íàäêðûëüå ñ ðèñóí-
êîì îïóøåíèÿ; 3 – óñèê; 4 – ïåðåäíÿÿ ÷àñòü òåëà, âèä ñáîêó; 5 – ïåðåäíåñïèíêà, âèä ñâåðõó (ñ
ðèñóíêîì îïóøåíèÿ); 6 – ïóíêòèðîâêà íà äèñêå ïåðåäíåñïèíêè; 7 – ïóíêòèðîâêà íà äèñêå íàä-
êðûëüåâ; 8 – ñåðåäèíà çàäíåãðóäè è 1-ãî âèäèìîãî áðþøíîãî ñòåðíèòà; 9 – ñðåäíÿÿ ëàïêà; 10 –
ýíäîôàëëóñ (âûâåðíóò). Ìàñøòàáíûå ëèíåéêè: 1,0 ìì (1); 0,5 ìì (2, 4, 5, 8); 0,2 ìì (3, 9); 0,1 ìì
(6, 7, 10).
Etymology. Species name comes from the Latin sparsus (sparse, diffuse, dis-
persed), reflecting diffused or spotted distribution of pubescence on the body.
Lasioderma tauricum sp. n. (fig. 5,1—11)
Material. Holotype {: Crimea, Karadag, 5.08.1991, meadow, net sweeping (I. T. A.). (“Lasioderma
kiesenwetteri Schils.” I. T. A. det., 1992) (ZMUM).
De s c r i p t i o n. Habitus as on fig. 5,1. Elytra brownish-yellow with yellow apices;
pronotum and head brownish; ventral surface black-brown; antennae (except brown
1st segment) and legs light yellow. Pubescence yellowish, homogenous, slightly raised.
He ad. Frons slightly convex. Eyes round, convex, separated by 1.7 eye diameters.
Antenna: 3rd segment almost as long as the 2nd one; 4—5th segments with concave
upper margin and convex lateral one; these two segments are of smaller size than next
6—10th segments; the latters have straight upper and lateral margins. 11th segment
about 3 times as long as wide and 1.9 times as long as the 10th segment (fig. 5,2). Last
segment of maxillary palpus is fusiform, about 3.5 times as long as its thickness, with
oblique truncate apex (fig. 5, 3).
P r ono t um 1.7 times as wide as long; anterior angles rectangular, posterior angles
obtuse, rounded. Pronotum convex more strongly near basal margin; sides not bulging
(fig. 5,4). Pubescence parted to two sides on disc (fig. 5, 5). Surface with fine, uniform
punctation, where punctures separated by 0.5—1.0 puncture diameter (fig. 5, 6).
S c u t e l l um semioval. Elytra 1.45 times as long as wide and 2.4 times as long as
pronotum. Surface with uniform punctation, punctures separated by 1—2 puncture
diameters (fig. 5, 7).
Me t a s t e r num about 1.5 times as long as the 1st visible abdominal sternite; ante-
rior rims approximated in the middle, 1st rim is straightened, the 2nd one slightly
curved forward (fig. 5, 8).
L e g s. Fore tibia not dilated to the apices. Tarsi as long as or longer than tibiae.
1st segment of hind tarsus 1.2 times as long as 2nd segment; 2nd segment 3 times as
long as 3rd segment; 3rd segment 1.2 times as long as 4th segment; 5th segment
1.7 times as short as 2nd segment (fig. 5, 9).
A ed e a gu s. Penis straight, almost parallel-sized, not narrowed to its apex; about
8 times as long as wide at apex; the latter has quadrate notch. Internal penial sac with
two longitudinal rows (4 + 9) of small spines, 5 small spines without arranging in rows,
and 1 large spine in apex. Penis is filled with thick, double “brush” in its two fifth of
basal part. Ends of parameres not deeply bifurcated, paramere outgrowth not reaching
paramere apex (fig. 5, 10). Cross-piece of genital ring segment very thin medially but
strongly widened to its sides (fig. 5, 11).
Length 2.4 mm, width 1.2 mm.
D i f f e r e n t i a l d i a g no s i s. The new species belongs to the group of yellow-
brown species with pubescence parted to two sides on pronotum and tibiae not widened
to their apices, which includes L. babadjanidesi, L kiesenwetteri, L. mikheechevi,
L. redtenbacheri, and L. sparsum sp. n. L. tauricum differs from L. babadjanidesi and
L. redtenbacheri by approximated anterior rims of metasternum (anterior rims evenly
rounded in those species), and from L. kiesenwetteri, L. mikheechevi, and L. sparsum
sp. n. first of all in shape of antennal segments: 4—5th segments are smaller and with
convex lateral margin whereas the next segments are larger and with straight lateral
margin; secondly, 1st tarsal segment almost as long as the 2nd segment and 5th seg-
ment is very short (1.7 times as short as the 2nd segment) in L. tauricum. In compared
species, only 4th segment has oblique upper margin, 5—10th segments more or less uni-
form, the 1st tarsal segment 1.5 times and more as long as the 2nd segment, and 5th
tarsal segment as long as the 2nd one in L. mikheechevi and L. sparsum and a little
203
shorter than the 2nd segment in L. kiesenwetteri. Difference in penial armaments is
shown in fig. 5, 10 and 4, 10 (also compare to: Toskina, 1999: fig. 44, 92, 140, and
Logvinovskij, 1985: fig. 228).
E t ymo l o g y. Species name is derived from the Latin geographic name Tauria
(Crimea), the type locality of the new species.
Key to species of the genus Lasioderma from Crimea and the Caucasus
Òàáëèöà äëÿ îïðåäåëåíèÿ âèäîâ ðîäà Lasioderma èç Êðûìà è Êàâêàçà
In this key, some data from the keys by Iablokoff-Khnzorian (1976), Logvinovskij
(1985), Toskina (1999), and articles and books by Arnoldi (1965), Español (1970,
1972, 1992), Zahradník (1998, 2000 et al.), and some others, were used.
1. Beetles black (excepting light margins of pronotum and elytra) or having something of black colour.
.............................................................................................................................................................. 2
- Beetles without black colour, but can be blackish-brown. .................................................................. 4
2. Antenna: 5—10th segments transverse, their lateral (i. e., lower) margins convex (Iablokoff-Khnzorian,
1976, fig. 1,10). Pronotum slightly campaniform, pubescence forms parting on pronotal disc; beetles
wholly black, pubescence grey (Roubal, 1916, S. 48). Length 2.5—3.4 mm. Czechoslovakia, Caucasus;
Crimea. ...................................................................................................... L. aterrimum Roubal, 1916.
204
Fig. 5. Lasioderma tauricum sp. n., holotype {: 1 – general outline of body; 2 – antenna; 3 – last segment
of maxillary palpus; 4 – anterior part of body, lateral view; 5 – pronotum, dorsal view (with pubescence
pattern); 6 – punctation on pronotal disc; 7 – punctation on elytral disc; 8 – the middle of metasternum
and 1st visible abdominal sternite; 9 – hind tibia and tarsus; 10 – aedeagus; 11 – distal part of genital ring.
Scale bar: 1.0 mm (1); 0.5 mm (4, 5, 8); 0.2 mm (2, 9—11); 0.1 mm (3, 6, 7).
Ðèñ. 5. Lasioderma tauricum sp. n., ãîëîòèï {: 1 – îáùèé êîíòóð òåëà; 2 – óñèê; 3 – ïîñëåäíèé ÷ëå-
íèê ÷åëþñòíûõ ùóïèêîâ; 4 – ïåðåäíÿÿ ÷àñòü òåëà, âèä ñáîêó; 5 – ïåðåäíåñïèíêà, âèä ñâåðõó (ñ
ðèñóíêîì îïóøåíèÿ); 6 – ïóíêòèðîâêà íà äèñêå ïåðåäíåñïèíêè; 7 – ïóíêòèðîâêà íà äèñêå íàä-
êðûëüåâ; 8 – ñåðåäèíà çàäíåãðóäè è 1-ãî âèäèìîãî áðþøíîãî ñòåðíèòà; 9 – çàäíèå ãîëåíü è ëàïêà;
10 – ýäåàãóñ; 11 – äèñòàëüíûé êîíåö ãåíèòàëüíîãî êîëüöà. Ìàñøòàáíûå ëèíåéêè: 1,0 ìì (1); 0,5 ìì
(4, 5, 8); 0,2 ìì (2, 9—11); 0,1 ìì (3, 6, 7).
- At least 8—10th ({) or 9—10th (}) antennal segments should be oblong .......................................... 3
3. Eyes shortly oval. Antenna: 5—7th ({) or 5—8th (}) segments only transverse, with convex lateral mar-
gin. Pronotum not campaniform, 1.7 times as wide as long; posterior angles developed, rounded;
pubescence parted to two sides on pronotal disc. Metasternum: anterior rims approximated in the mid-
dle, 1st rim straightened, the 2nd one curved forwards. Internal penial sac with 3 rows (7 + 5 + 11)
of small spines. Pronotal and elytral apical margins reddish; sometimes elytra dark brown. Pubescence
homogenous, yellowish-grey, appressed. Length 2.4—4.5 mm. Steppes of Southern Europe (Arnoldi,
1965), Crimea, Caucasus. ........................................................................ L. obscurum (Solsky, 1867).
- Eyes round. Antenna: 6—10th segments oblong, with straight lateral margin. Pronotum 1.5 times as wide
as long, its posterior angles slightly developed; pubescence not parted to two sides. Metasternum: ante-
rior rims approximated and straightened in the middle. Internal penial sac with 1 row of 6—7 small
spines and 3 spines of median size. Length 2.25 mm, width 1.1 mm. Armenia. ..................................
.................................................................................................................................... L. morulum sp. n.
4. Pronotal surface with very large punctation, punctures are seen with magnification x10. Internal penial
sac with 2 rows (8 + 10) of small spines and 4 large spines (Logvinovskij, 1985, fig. 233). Beetles rufous.
Length 1.2—2.1 mm. Crimea (Arnoldi, 1965); Israel (Logvinovskij, 1985; Zahradník et al., 2000). ....
.................................................................................................................. L. punctulatum Reitter, 1884.
- Surface with much smaller punctation. .............................................................................................. 5
5. Pubescence not parted to two sides on pronotal disc. ........................................................................ 6
- Pubescence parted to two sides on pronotal disc. ............................................................................ 12
6. Fore tibiae widened to their apices. .................................................................................................... 7
- Fore tibiae not widened to their apices. .............................................................................................. 8
7. Entirely blackish-brown. Antenna: 4—5th segments with concave upper and convex lateral margin;
6—9th segments transverse, with straight upper and lateral margins. Pronotum with clear posterior
angles. Metasternum: anterior rims approximated in the middle, 1st rim straightened, the 2nd one
curved forwards. 1st segment of hind tarsus 3 times as long as the 2nd one. Internal penial sac with
“brush” in its apical half, basal ends of this “brush” are finished with 3 small needle-shaped sclerites.
Pubescence dark grey, homogenous, appressed. Length 3.2 mm, width 1.75 mm. Northern Caucasus.
.................................................................................................................................... L. prolixum sp. n.
- Brownish-red, antennae light yellow. Antenna: 4—10th segment cup-shaped. Pronotum without poste-
rior angles. Metasternum: anterior rims evenly rounded. 1st segment of hind tarsus twice as long as the
2nd one. Internal penial sac with 7 spines of different sizes (Español, 1972, fig. 33; 1992, fig. 58B).
Length 1.5—2.9 mm. Cosmopolite. ........................................................ L. serricorne (Fabricius, 1792)
8. Eyes very large: separated by 1.35 eye diameters. Antenna: 3rd segment 1.5 times as long as the 2nd
one; 4—10th segments with convex lateral margin, the segments oblong except 5—6th segments.
Pronotum 1.5 times as wide as long, posterior angles developed, rounded. Elytra 1.5 times as long as
wide. Metasternum: anterior rims approximated and straightened in the middle. Internal penial sac with
2 rows (9 + 5) of small spines, 2 large spines, and 2 baculiform sclerites (Toskina, 1999, fig. 126).
Beetle reddish-brown, antennae lighter. Pubescence golden-yellow, heterogeneous, partly inclined, long
inclined hairs not arranged in rows. Length 2.3 mm, width 1.6 mm. Azerbaijan. ................................
...................................................................................................................... L. oculeum Toskina, 1999.
- Eyes smaller ........................................................................................................................................ 9
9. Antenna: 3rd segment about as long as the 2nd one. ........................................................................ 10
- 3rd antennal segment 1.5 times as long as the 2nd one; 5—10th segments with convex lateral one
(Español, 1970, fig. 17). Pronotum 1.4 times as wide as long; posterior angles developed. Elytra 1.6
times as long as wide. Metasternum: anterior rims approximated in the middle and curved forwards.
Internal penial sac with 14 small spines, 2 spines of medium sizes, and 2 very large spines (Español,
1992, fig. 58E). Dorsal surface dark brown, elytral apex or elytra as a whole are red. Pubescence
homogenous. Length 2.0—3.2 mm. Black Sea steppe (leg. Arnoldi, coll. ZISP); Crimea, Caucasus
(Logvinovskij, 1985); Mediterranea (Español, 1992); Turkey, Canary Is. (Zahradník, 1998); Israel
(Zahradník et al., 2000). .................................................................... L. haemorrhoidale (Illiger, 1807)
10. Pubescence homogenous and appressed on elytra. Antenna: 4th segment with oblique and concave
upper margin and straight lateral one. 5—10th segments with convex lateral margin. Pronotum 1.5—1.6
times as wide as long, posterior angles not developed. Elytra 1.8—2.0 times as long as wide; surface
wrinkled. Metasternum: anterior rims evenly rounded. Beetles greyish-yellow. Length 4.5—5.0 mm,
width 1.8—1.9 mm. Crimea. .......................................................................................... L. dolini sp. n.
- Elytra with dual pubescence. ............................................................................................................ 11
11. Legs: 1st segment of hind tarsus 1.5 times as long as the 2nd one. Antenna: middle segments 1.6—1.7
times as long as wide (Logvinovskij, 1985, fig. 209). Pronotum: posterior angles rounded. Beetles yel-
low-brown, antennae dark brown. Length 3.2—4.2 mm. Azerbaijan. ....................................................
...................................................................................................... L. kryzhanovskii Logvinovskij, 1977.
- Legs: 1st segment of hind tarsus is as long as the 2nd one. Antenna: middle segments hardly oblong
(Toskina, 1999, fig. 71). Pronotum 1.6 times as wide as long, posterior angles slightly developed. Elytra
1.5 times as long as wide. Metasternum: anterior rims approximated and straightened in the middle.
205
Dorsal surface reddish-brown, ventral surface darker, antennae and legs yellow. Length 2.8 mm, width
1.4 mm. Armenia. .................................................................................... L. mazokhini Toskina, 1999.
12. Anterior rims of metasternum `are evenly rounded in the middle. .................................................. 13
- Anterior rims of metasternum are approximated in the middle. ...................................................... 15
13. Antenna: 3rd segment 1.5—1.7 times as long as the 2nd one; 4—10th segments with straight lateral mar-
gins. Pronotum 1.44 times as wide as long; its posterior angles are developed; pubescence parted to two
sides on apical third of pronotum. Elytra 1.7 times as long as wide; surface with uniform punctation,
punctures not merged in wrinkles. 1st segment of hind tarsus 1.5 times as long as the 2nd one. Beetles
brick-red. Pubescence heterogeneous. Length 3.9—5.1 mm. Steppe and partially-wooded steppe in
South Russia (Arnoldi, 1965); Crimea, the Caucasus, Kazakhstan (Logvinovskij, 1985); the Middle and
South Europe, North Africa (Español, 1992); Turkey (Zahradník, 1998); Israel (Zahradník et al.,
2000). .................................................................................................... L. redtenbacheri (Bach, 1852).
- Antenna: 3rd segment almost as long as the 2nd one ...................................................................... 14
14. Pronotum 1.8 times as wide as long; posterior angles are missing. Pubescence forming wave-picture on
pronotum. Elytra 1.5—1.6 times as long as wide. 1st tarsal segment of hind tarsus twice as long as the
2nd one. Internal penial sac with 8 rather large spines (Toskina, 1999, fig. 44). Beetles dark greyish-
brown, pubescence homogenous, yellowish-brown, and appressed. Length 2.8—3.5 mm. Transcaucasus.
................................................................................................................ L. babadjanidesi Reitter, 1913.
- Pronotum 1.6—1.7 times as wide as long; posterior angles slightly developed. Elytra 1.5 times as long
as wide. 1st segment of hind tarsus 1.5 times as long as the 2nd one. Internal penial sac with row of 3
small spines and 5 spines of medium size. Beetles from dark brown to reddish-brown, apical third of
elytra greyish-yellow. Pubescence homogenous, appressed, golden-grey, not waving on pronotum.
Length 2.7—3.2 mm, width 1.5—1.6 mm. Azerbaijan. .......................... L. mikheechevi Toskina, 1999.
15. Pubescence parted to two sides near pronotal apical margin only; posterior angles developed and round-
ed. Elytra 1.56 times as long as wide, surface with uniform punctation. Internal penial sac with 3 rows
(5 + 5 + 14) of small spines and 2 large spines. Beetles yellow-red colour, antennae and legs yellow.
Pubescence light yellow, homogenous, dense, raised. Length 2.8—3.5 mm. Crimea (Arnoldi, 1965);
Crimea, the Caucasus, West Kazakhstan (Logvinovskij, 1985); South Europe, Turkey (Zahradnik,
1998); Israel (Zahradnik et. al., 2000). ................................................ L. kiesenwetteri Schilsky, 1899
- Pubescence parted to two sides all over the disc of pronotum .......................................................... 16
16. Pubescence unevenly arranged, maculate on elytra. Antenna: 4th segment with oblique and straight
upper margin, 5—10th segments with slightly convex lateral margin. Pronotum 1.55 times as wide as
long, posterior angles slightly developed. Elytra 1.6 times as long as wide, surface with dual punctation.
1st segment of hind tarsus 1.5 times as long as the 2nd one. Internal penial sac with 1 row of 8 small
spines, 2 detached small spines, and 3 large spines. Beetle wholly brown, pubescence heterogeneous,
raised. Length 3.5 mm, width 1.65 mm. Northern Caucasus. .................................... L. sparsum sp. n.
- Surface of elytra with homogenous pubescence. Antenna: 4—5th segments with oblique and concave
upper margin and convex lateral one; these segments are of smaller size than next segments; 6—10th
segments with straight lateral margin. Pronotum 1.7 times as wide as long, posterior angles developed.
Elytra 1.45 times as long as wide, surface with uniform punctation. Metasternum: 1st rim straightened,
2nd curved forwards. 1st segment of hind tarsus nearly as long as the 2nd one. Internal penial sac with
2 rows (14 + 9) of small spines, 5 small spines not arranged in rows, and 1 large spine. Beetle brown-
ish-yellow, head brown. Pubescence homogenous, slightly raised. Length 2.4 mm, width 1.2 mm.
Crimea ...................................................................................................................... L. tauricum sp. n.
Discussion
Lohse considered L. aterrimum Roubal as synonym of L. obscurum (Solsky)
(Lohse, 1969: 54), but Iablokoff-Khnzorian figured parts of L. aterrimum holotype
(body outline, antennae, parts of aedeagus) and separated these two species in his key
(Iablokoff-khnzorian, 1976, fig. 8, 10, 5; p. 126). Zahradník when describing a new
species L. anatolica from Turkey compared it with other species of black colour, includ-
ing L. aterrimum in the differential diagnosis, so this author admitted L. aterrimum as
a separate species (Zahradník, 1996: 98). We included this species in the key.
We are unable to recognize and include L. redtenbacheri var. caucasicum Pic, 1904
in the key because its holotype was unavailable in this study, and author’s description
of this form is very short (beetle of clay-rufousish colour, with very dense pubescence).
There is a specimen in the ZISP collection which was identified as L. obscurum var cau-
casicum Pic. A. Reichardt determinated it as a valid species L. caucasicum Pic. L. obscu-
rum and L. caucasicum Pic – Reichardt differ rather strongly in metasternum struc-
tures, in eyes, at last in beetles colour (anterior rims evenly rounded, eyes round, bee-
206
tle with elytra black only in “caucasicum” whereas anterior rims of metasternum are
approximated very strongly, eyes oval, beetles usually wholly black in L. obscurum). We
do not know if A. Reichardt had ever seen the holotype of L. redtenbacheri var. cauca-
sicum Pic. Short description L. redtenbacheri var. caucasicum by Pic is more related to
L redtenbacheri as its variant than to L. obscurum. We leave this question open for the
present time.
The author is sincerely grateful to Dr. O. Merkl (HMNH) for providing with type material,
E. P. Nartshuk and S. V. Andreeva (ZISP), N. B. Nikitsky (ZMUM), and N. L. Klepikova (Moscow) for
great assistance in this work.
Arnoldi L. V. The family Anobiidae// Opredelitel’nasekomykh evropeiskoj chasti SSSR. 2. Coleoptera and
Strepsiptera / Ed. G. Ya. Bei-Bienko. – M. ; L. : Nauka Publ., 1965. – Vol. 2. – P. 244—257. –
Russian : Àðíîëüäè Ë. Â. Ñåì. Anobiidae // Îïðåäåëèòåëü íàñåêîìûõ åâðîïåéñêîé ÷àñòè ÑÑÑÐ.
Æåñòêîêðûëûå è âååðîêðûëûå.
Español F. Ergebnisse der zoologischen Forschungen von Dr. Z. Kaszab in der Mongolei N 249 (Col.
Anobiidae) // EOS (Revista española entomologia). – 1970. – 46. – P. 41—56.
Español F. Contribution al conocimiento de los Xyletininae (Col., Anobiidae) del Gabόn (Mision H. Coiffait,
1963) // Biologia Gabonica, revue. – 1972. – 8, N 2. – P. 175—189.
Español F. Coleoptera, Anobiidae // Fauna Iberica. – Madrid : Museo Nacional de Ciencias Naturales,
CSIC, 1992. – Vol. 2. – 195 p.
Iablokoff-Khnzorian S. M. New species of beetle-borer from Darvaz // Doklady Academii Nauk ArmSSR. –
1976. – 63, N 2. – P. 124—127. – Russian: ßáëîêîâ-Õíçîðÿí Ñ. Ì. Íîâûå âèäû æóêîâ-òî÷èëüùè-
êîâ èç Äàðâàçà // Äîêë. ÀÍ ÀðìÑÑÐ.
Logvinovskij V. D. Wood-borers the family Anobiidae. – Leningrad : Nauka Publ., 1985. – 175 p. – (Fauna
USSR. New Ser. N 131. Coleoptera, Vol. 14, Fasc. 2). – Russian :
Ëîãâèíîâñêèé Â. Ä. Òî÷èëüùèêè – ñåì. Anobiidae. – Ë. : Íàóêà, 1985. – (Ôàóíà ÑÑÑÐ. Ñåð.
Íîâ., ¹ 131. Íàñåêîìûå æåñòêîêðûëûå ; Ò. 14, âûï. 2).
Lohse G. A. Anobiidae // Die Käfer Mitteleuropas / H. Freude, K. W. Harde, G. A. Lohse. – Krefeld :
Goecke & Evers, 1969. – 8. – S. 27—59.
Pic M. Diagnoses de Coléoptères palearctiques et exotiques // L’Echange. – 1904. – 20. – P. 33—38.
Roubal J. Neue palaearktischen Coleopteren // Coleopterol. Rundschau. – 1916. – 5, H. 4/6. – S. 45—48.
Toskina I. N. New and rare species of the genus Lasioderma Stephens, 1835 (Coleoptera: Anobiidae) //
Russian Entom. J. – 1999. – Vol. 8, N 1. – P. 23—46.
Zahradník P. Fore new species of Anobiidae (Coleoptera) from Turkey and Rhodes Island // Folia
Heyrovskyana. – 1996. – 4, N 3. – P. 93—101.
Zahradník P. Anobiidae of Turkey (Coleoptera) // Klapalekiana. – 1998. – 34. – P. 263—286.
Zahradník P., Chikatunov V., Pavlíček T. Catalogue of beetles (Coleoptera) of Israel and adjacent areas: 2.
Anobiidae // Klapalekiana. – 2000. – 36. – P. 307—319.
207
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| id | nasplib_isofts_kiev_ua-123456789-65800 |
| institution | Digital Library of Periodicals of National Academy of Sciences of Ukraine |
| issn | 0084-5604 |
| language | English |
| last_indexed | 2025-12-07T18:01:06Z |
| publishDate | 2011 |
| publisher | Інститут зоології ім. І.І. Шмальгаузена НАН України |
| record_format | dspace |
| spelling | Toskina, I.N. 2014-07-02T19:09:39Z 2014-07-02T19:09:39Z 2011 New Species of the Genus Lasioderma (Coleoptera, Anobiidae) from Crimea and the Caucasus / I.N. Toskina // Вестник зоологии. — 2011. — Т. 45, № 3. — С. 195–207. — Бібліогр.: 13 назв. — англ. 0084-5604 https://nasplib.isofts.kiev.ua/handle/123456789/65800 395.766.4(47) Five species of the genus Lasioderma Stephens, 1835 from Crimea (L. dolini sp. n., L. tauricum sp. n.), North Caucasus (L. prolixum sp. n., L. sparsum sp. n.), and Armenia (L. morulum sp. n.) are described. A key to 17 species of the genus Lasioderma occurring in these regions is compiled. Описаны 5 видов рода Lasioderma Stephens, 1835 из Крыма (L. dolini sp. n., L. tauricum sp. n.), Северного Кавказа (L. prolixum sp. n., L. sparsum sp. n.) и Армении (L. morulum sp. n.). Составлена определительная таблица 17 видов рода Lasioderma для названного региона. The author is sincerely grateful to Dr. O. Merkl (HMNH) for providing with type material, E. P. Nartshuk and S. V. Andreeva (ZISP), N. B. Nikitsky (ZMUM), and N. L. Klepikova (Moscow) for great assistance in this work. en Інститут зоології ім. І.І. Шмальгаузена НАН України Вестник зоологии Фауна и систематика New Species of the Genus Lasioderma (Coleoptera, Anobiidae) from Crimea and the Caucasus Новые виды рода Lasioderma (Coleoptera, Anobiidae) из Крыма и Кавказа Article published earlier |
| spellingShingle | New Species of the Genus Lasioderma (Coleoptera, Anobiidae) from Crimea and the Caucasus Toskina, I.N. Фауна и систематика |
| title | New Species of the Genus Lasioderma (Coleoptera, Anobiidae) from Crimea and the Caucasus |
| title_alt | Новые виды рода Lasioderma (Coleoptera, Anobiidae) из Крыма и Кавказа |
| title_full | New Species of the Genus Lasioderma (Coleoptera, Anobiidae) from Crimea and the Caucasus |
| title_fullStr | New Species of the Genus Lasioderma (Coleoptera, Anobiidae) from Crimea and the Caucasus |
| title_full_unstemmed | New Species of the Genus Lasioderma (Coleoptera, Anobiidae) from Crimea and the Caucasus |
| title_short | New Species of the Genus Lasioderma (Coleoptera, Anobiidae) from Crimea and the Caucasus |
| title_sort | new species of the genus lasioderma (coleoptera, anobiidae) from crimea and the caucasus |
| topic | Фауна и систематика |
| topic_facet | Фауна и систематика |
| url | https://nasplib.isofts.kiev.ua/handle/123456789/65800 |
| work_keys_str_mv | AT toskinain newspeciesofthegenuslasiodermacoleopteraanobiidaefromcrimeaandthecaucasus AT toskinain novyevidyrodalasiodermacoleopteraanobiidaeizkrymaikavkaza |