A revision of afrotropical species of the eupyrgota (diptera, pyrgotidae): the spinifemur group and latipennis subgroup of species
Based on the study of the type specimens described
 by F. Walker, F. Hendel, E. Brunetti and P. Vanschuytbroeck and exhaustive material from many
 entomological collections, species of the spinifemur group and latipennis subgroup of the genus
 Eupyrgota Coquillett...
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| Datum: | 2006 |
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Інститут зоології ім. І.І. Шмальгаузена НАН України
2006
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| Zitieren: | A revision of afrotropical species of the eupyrgota (diptera, pyrgotidae): the spinifemur group and latipennis subgroup of species / V.A. Korneyev // Вестн. зоологии. — 2006. — Т. 40, № 1. — С. 3-25. — Бібліогр.: 18 назв. — англ. |
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Digital Library of Periodicals of National Academy of Sciences of Ukraine| _version_ | 1860148689265229824 |
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| author | Korneyev, V.A. |
| author_facet | Korneyev, V.A. |
| citation_txt | A revision of afrotropical species of the eupyrgota (diptera, pyrgotidae): the spinifemur group and latipennis subgroup of species / V.A. Korneyev // Вестн. зоологии. — 2006. — Т. 40, № 1. — С. 3-25. — Бібліогр.: 18 назв. — англ. |
| collection | DSpace DC |
| description | Based on the study of the type specimens described
by F. Walker, F. Hendel, E. Brunetti and P. Vanschuytbroeck and exhaustive material from many
entomological collections, species of the spinifemur group and latipennis subgroup of the genus
Eupyrgota Coquillett occurring in the Afrotropical Region and Arabian Peninsula are redescribed. The
following synonymy is established. Eupyrgota latipennis (Walker, 1849), comb. n. (= Oxycephala
latipennis Walker) = Adapsilia ypsilon Hendel, 1914, syn. n. Eupyrgota spinifemur (Hendel, 1934) comb. n.
(= Adapsilia spinifemur Hendel) = Metropina nigra Vanschuytbroeck, 1963, syn. n. Eupyrgota saegeri
(Vanschuytbroeck, 1963) comb. n. is transferred from Lygiohypotyphla Hendel.
На основании изучения типовых материалов, описанных Ф. Уокером, Ф. Генделем, Ю. Брунетти и П. Вансхётбруком и исчерпывающих материалов из многочисленных энтомологических коллекций, представлены описания видов из группы spinifemur и подгруппы latipennis рода Eupyrgota, встречающихся в Афротропической области и на Аравийском полуострове. Установлена следующая синонимия. Eupyrgota latipennis (Walker, 1849), comb. n. (= Oxycephala latipennis Walker) = Adapsilia ypsilon Hendel, 1914, syn. n. Eupyrgota spinifemur (Hendel, 1934) comb. n. (= Adapsilia spinifemur Hendel) = Metropina nigra Vanschuy-tbroeck, 1963, syn. n. Eupyrgota saegeri (Vanschuytbroeck, 1963) comb. n. перемещена из Lygiohypo-typhla Hendel.
|
| first_indexed | 2025-12-07T17:51:26Z |
| format | Article |
| fulltext |
UDC 595.773.4
A REVISION OF AFROTROPICAL SPECIES
OF THE EUPYRGOTA (DIPTERA, PYRGOTIDAE):
THE SPINIFEMUR GROUP AND
LATIPENNIS SUBGROUP OF SPECIES
V. A. Korneyev
Royal Museum for Central Africa,
Leuvensesteenweg 13, Tervuren, B-3080 Belgium and
Schmalhausen Institute of Zoology, NAS of Ukraine,
Bogdan Chmielnicky str., 15, Kyiv, 01601 Ukraine
E-mail: kot-vasily@yandex.ru
Accepted 24 October 2005
Ðåâèçèÿ àôðîòðîïè÷åñêèõ âèäîâ ðîäà Eupyrgota (Diptera, Pyrgotidae): ãðóïïà âèäîâ spinifemur è ïîä-
ãðóïïà latipennis. Êîðíååâ Â. À. — Íà îñíîâàíèè èçó÷åíèÿ òèïîâûõ ìàòåðèàëîâ, îïèñàííûõ
Ô. Óîêåðîì, Ô. Ãåíäåëåì, Þ. Áðóíåòòè è Ï. Âàíñõ¸òáðóêîì è èñ÷åðïûâàþùèõ ìàòåðèàëîâ èç
ìíîãî÷èñëåííûõ ýíòîìîëîãè÷åñêèõ êîëëåêöèé, ïðåäñòàâëåíû îïèñàíèÿ âèäîâ èç ãðóïïû
spinifemur è ïîäãðóïïû latipennis ðîäà Eupyrgota, âñòðå÷àþùèõñÿ â Àôðîòðîïè÷åñêîé îáëàñòè è
íà Àðàâèéñêîì ïîëóîñòðîâå. Óñòàíîâëåíà ñëåäóþùàÿ ñèíîíèìèÿ. Eupyrgota latipennis (Walker,
1849), comb. n. (= Oxycephala latipennis Walker) = Adapsilia ypsilon Hendel, 1914, syn. n. Eupyrgota
spinifemur (Hendel, 1934) comb. n. (= Adapsilia spinifemur Hendel) = Metropina nigra Vanschuy-
tbroeck, 1963, syn. n. Eupyrgota saegeri (Vanschuytbroeck, 1963) comb. n. ïåðåìåùåíà èç Lygiohypo-
typhla Hendel.
Êëþ÷åâûå ñ ëîâ à: Diptera, Cyclorrhapha, Tephritoidea, Pyrgotidae, Eupyrgota, Àôðîòðîïè÷åñ-
êàÿ îáëàñòü, ôàóíà, òàêñîíîìèÿ, ñèíîíèìèÿ.
A Revision of Afrotropical Species of the Eupyrgota (Diptera, Pyrgotidae): the spinifemur Group and
latipennis Subgroup of Species. Korneyev V. A. — Based on the study of the type specimens described
by F. Walker, F. Hendel, E. Brunetti and P. Vanschuytbroeck and exhaustive material from many
entomological collections, species of the spinifemur group and latipennis subgroup of the genus
Eupyrgota Coquillett occurring in the Afrotropical Region and Arabian Peninsula are redescribed. The
following synonymy is established. Eupyrgota latipennis (Walker, 1849), comb. n. (= Oxycephala
latipennis Walker) = Adapsilia ypsilon Hendel, 1914, syn. n. Eupyrgota spinifemur (Hendel, 1934) comb. n.
(= Adapsilia spinifemur Hendel) = Metropina nigra Vanschuytbroeck, 1963, syn. n. Eupyrgota saegeri
(Vanschuytbroeck, 1963) comb. n. is transferred from Lygiohypotyphla Hendel.
Ke y wo r d s: Diptera, Cyclorrhapha, Tephritoidea, Pyrgotidae, Eupyrgota, Afrotropical Region, fauna,
taxonomy, new taxa, synonymy.
Pyrgotidae are medium to large-sized (4–18 mm) acalyptrate flies, which usually can be recognized by
pictured wings with short lobate cell bcu, rather slender body, oblique face, and stiletto-like aculeus that is much
shorter than the oviscape. The World fauna includes about 365 valid species names in ca. 55 genera, with the
greatest number of species in the tropics (Korneyev, unpublished data). Up to now, over 148 nominal species
assigned to 24 genera, or almost one half of the described species of the family, were recorded from the
Afrotropical Region (Steyskal, 1980). However, no comprehensive taxonomic revisions have been provided for
the pyrgotids from the Oriental and Afrotropical Regions. Most paleotropical species are inadequately described
and usually have not been figured, so they can be recognized only by comparison with type specimens.
M. Aczél (1956 a, b, c) and recently S.-K. Kim and H.-Y. Han (2000, 2001) and V. A. Korneyev
(2004) elaborated the most comprehensive approach to diagnostics of the Pyrgotidae, which takes into con-
sideration numerous morphological (including genitalic) characters, not used in description of Afrotropical
pyrgotids.
Vestnik zoologii, 40(1): 3–25, 2006
© V. A. Korneyev, 2006
Ôàóíà è ñèñòåìàòèêà
While preparing the Catalogue of the Pyrgotidae of the World, material from the Afrotropical Region
deposited in several African, European and American collections was examined. This study has shown that
the family needs a thorough taxonomic revision based on studies of exhaustive material. This paper is the first
in a series of publications resulting from study supported by the Belgian Federal Office for Scientific,
Technical and Cultural Affairs Research Fellowship and conducted in the Royal Museum for Central Africa,
Tervuren, Belgium in 2005–2006.
The present paper is divided into two parts. The first part concerns the spinifemur group of species, and
the latipennis subgroup of the latipennis group of species. A revision of the varipennis and melancholica
subgroups of the latipennis group of species, including descriptions of several new species and a key to
Afrotropical species of the genus Eupyrgota shall be published in the next issue of this journal.
Material
The specimens examined in this study are deposited in the following collections: AMGS — Albany
Museum, Grahamstown, South Africa; AMNH — American Museum of Natural History, New York, U.S.A.;
BMNH — the Natural History Museum, London, U.K.; CMNH — the Carnegie Museum of Natural
History, Pittsburgh, U.S.A.; DEI — Deutsche Entomologisches Institut, Mu�ncheberg, Germany; FBUB —
Fakulta �t fu �r Biologie, Universita �t Bielefeld, Germany; HMNH – Hungarian Museum of Natural History,
Budapest; RBINH — Royal Belgian Institute of Natural History, Brussels, Belgium; RMCA — Royal
Museum of Central Africa, Tervuren, Belgium; NHMW — Naturhistorisches Museum Wien, Austria;
NMKE — National Museums of Kenya, Nairobi; NMPM — the Natal Museum, Pietermaritzburg, South
Africa; RNHL — Nationaal Natuurhistorisch Museum, Leiden, the Netherlands; SANC — South African
National Collection of Insects, Pretoria; SIZK — Schmalhausen Institute of Zoology, Kyiv, Ukraine;
SMNS — Staatliches Museum fu �r Naturkunde Stuttgart, Germany; USNM — National Museum of Natural
History, Smithsonian Instituion, Washington, D. C., U.S.A.; ZMAN — Zoologisch Museum, Instituut voor
Taxonomische Zoologie, Universiteit van Amsterdam, the Netherlands; ZMHB — Museum fu�r Naturkunde
der Humboldt-Universita�t zu Berlin, Germany, ZSSM — Zoologische Staatsammlung Mu �nchen, Germany.
The slash character (/) is used to separate lines, and the square brackets are for deciphered
abbreviations in the literally cited labels. The non-type material is arranged alphabetically by country names,
then from the West to the East and from the North to the South within each country; and finally, by the
year, month and day of collecting; the collector(s) name(s) and the abbreviation of the depositary enclose
the list of non-type specimens and are provided only once in the end if repeated.
Copyrights of the pictures of specimens deposited in the BMNH used in this paper belong to the
Natural History Museum, London.
Classification of Pyrgotidae
The most detailed classification of the family was proposed by M. Aczél (1956 a,
b, c), and modified by D. K. McAlpine (1990); it is subdivided into two subfamilies,
Teretrurinae (two South American genera with four species) and Pyrgotinae, which
includes the vast majority of genera and species. The Pyrgotinae are divided into two
tribes: Toxurini from the Australasian Region, with more than 60 described species in at
least six genera (tribal position of some genera is uncertain), and Pyrgotini with more
than 275 species in 35–40 genera from all zoogeographical regions. The subfamily Loch-
mostylinae is now considered to be the family Ctenostylidae (McAlpine, 1990).
G. C. Steyskal (1972) suggested that details of the structure of the postabdomen can pro-
vide a firm classification of the family. Recently, S.-K. Kim and H.-Y. Han (2000, 2001)
and then V. A. Korneyev (2004) figured and described terminalia of Eastern Palaearctic
pyrgotids. The further study of terminalia of Pyrgotidae (Korneyev, De Coninck, in
prep.; Korneyev, in prep.) has the following taxonomic and phylogenetic implications.
In all examined Pyrgotini, the epandrium and lateral surstylus are completely
devoid of setulae (at most, short setulae reminiscent of trichoid sensilla are developed
on the surstyli). The Afrotropical genus Toxopyrgota Hendel, a few Australian genera of
Toxurini that were studied (Cardiacera Macquart and Neotoxura Hendel), and
Teretrurinae have a setose epandrium, as in other Tephritoidea. The absence of epan-
drial setulae is evidently universal for the Palaearctic, Oriental, Afrotropical and New
World Pyrgotini. Another character, which is universal for all examined Palaearctic,
Oriental and Afrotropical species of this tribe, is the presence of the subocular dilation
of the parafacial, a sclerite below the ventral margin of the compound eye, separated
4 V. A. Korneyev
from the gena and occiput by a soft, non-sclerotized area, subdivided into the genal
groove and narrow postocular groove (the latter character is apparently universal for all
Pyrgotidae). All Teretrurinae, Toxurini, Toxopyrgota incertae sedis, Old World Tephri-
topyrgota Hendel, Tephritocampylocera Hendel, Pyrgotina Malloch (Pyrgotinae) and all
New World species of Pyrgotini do not possess the subocular dilation, and the parafa-
cial is gradually narrowed below the ventral side of the eye. In all examined Pyrgotinae
the subgenae (narrow sclerotized stripes ventral to antennal grooves and anterior to the
vibrissal edge and fronto-genal suture) are connected by the supraclypeal sclerite (meso-
facial plate sensu Acze´l, 1956 a), a bridge formed by a sclerotized portion of the fron-
to-clypeal membrane that has a triangular or pentagonal shape and is joined to the
facial carina dorsally. In Toxopyrgota, a similar structure is membranous, and in
Teretrurinae it is primarily absent, as the shape of face is entirely different (Korneyev,
unpublished data).
The presence of the subocular dilation is a synapomorphy of several Old World
genera (Adapsilia Waga, Campylocera Macquart, Euphya Wulp, Eupyrgota Coquillett,
Pyrgotomyia Hendel, Siridapha Enderlein and some other minor genera related to
them). If this group is considered a separate subtribe or tribe, a name should be derived
from the available family-group name Adapsilioidi Rondani, 1869, which is considered
now to be a synonym of Pyrgotini.
In this paper, I generally follow the concept of Eupyrgota Coquillett proposed by
S.-K. Kim and H.-Y. Han (2001). Morphological terminology for the parts of the
female terminalia follows White et al. (1999), rather than G. C. Steyskal (1972). The
tergosternite 7 forms a large oviscape; its antero-ventral apodeme is often an unusually
large flap reaching the dorsal side, connected by strong longitudinal muscles with the
walls of oviscape and able to close up the whole anterior opening of the oviscape. In
the distal, or posterior margin of the oviscape bordering the eversible membrane, some
sclerotized lobes can be recognized, including two medio-apical lobes, the ventral (ven-
tro-medial lobe) and often the dorsal (dorso-medial lobe) are extended onto the
eversible membrane; at rest, they are inverted inside the oviscape. Each of two lateral
lobes, symmetrical and usually developed on the ventral side, as a rule consists of a
darker and softer ventral field of mechanoreceptive sensilla, visible even when the mem-
brane is completely inverted, and of an elongate, taenia-like stripe, which extends on
the membrane latero-ventrally and bears no sensilla. In Eupyrgota, there is a pair of dor-
sal fields of mechanoreceptive sensilla lateral to the dorso-medial lobe. The eversible
membrane in some cases has paired fields of short acanthi forming a rasper-like struc-
ture posterior and between the ventro-medial and lateral lobes; from their topology,
these structures may be derivatives of taeniae as seen in other Tephritoidea. In
Eupyrgota they are modified into paired hooks (ovipositungui sensu Steyskal, 1972). The
aculeus is unusually short compared to the oviscape, and very acute, with the cercal
unit completely integrated into its apex without a trace of fusion, and the 8th sternite
(or the ventral lobes) narrow and rather soft.
Eupyrgota Coquillett, 1898
Typ e s p e c i e s: Eupyrgota luteola Coquillett, 1898, by original designation.
D i a gno s i s. Species of Eupyrgota can be recognized from the other Old World
Pyrgotini by combination of the following characters: presternum bearing a pair of fin-
ger-like lobes, femora apico-ventrally with postero- and antero-ventral row of thickened
setae (fig. 3, 4; 4, 10, 13; 7, 11) and the oviscape with a pair of heavily sclerotized
hook-like lobes at apico-ventral margin (fig. 2, 4; 5, 5; 8, 1–2). The presence of fin-
ger-like lobes of the presternum is the unique character of Eupyrgota, which does not
occur in other pyrgotid genera and supports its monophyly. The rows of spurious setae
on the femora are known also in Euphya Wulp (Pyrgotini), Epice Paramonov
5A Revision of Afrotropical Species of the Eupyrgota (Diptera, Pyrgotidae)...
(Toxurini) and numerous taxa in the families Richardiidae, Tephritidae and Platysto-
matidae. Furthermore, in a few Oriental species of Eupyrgota they are poorly expressed.
Species of both Euphya and Epice can be recognized by the saddle-like shape of the
presternum without prominent lobes and the oviscape without hook-like lobes. Females
of East Asian (Palaearctic and possibly Oriental) species Adapsilia hirtoscutellata Hendel
and A. myopoides Chen both have similarly sclerotized hook-like lobes of the oviscape,
which have somewhat different position and are separated by a small sclerotized ven-
tro-medial lobe or sclerite (Korneyev, 2004; Nartshuk, Korneyev, 2005: fig. 13–14).
These species can be distinguished from Eupyrgota by the absence of finger-like lobes
of the presternum and spur-like setae on the femora.
De s c r i p t i o n. Moderate or large (wing length 5.0–18.0 mm) wasp-like flies,
often with moderately petiolate abdomen, yellow and brown coloration of body and
wings with darkened band or two brown spots in anterior half. Head higher than wide,
short or moderately long setose; ocelli absent; parafacial very short setulose, with sub-
ocular dilation; facial carina developed or absent; supraclypeal sclerite low to moder-
arely high (at most 0.33 times as high as face); lateral vertical seta short or indistin-
guishable; antenna as long as face, flagellomere 1 almost as long as pedicel; palpus usu-
ally as long as flagellomere 1 (or slightly longer in some Oriental species), parallel-sided
or apically expanded; labellum moderately large, fleshy; presternum bearing a pair of
finger-like lobes; presutural supra-alar and prescutellar acrostichal setae absent; scutel-
lum with 1–5 pairs of setae; wing with humeral and subcostal break present; costa
reaching medial vein; Sc straight, narrowly broken before costa; R2+3 with spurious
vein, except in few Oriental species; cell bm and bcu almost entirely bare, with faint
yellowish tinge; Cu2 short; halter yellow; midcoxa anteroventrally setose and setulose
but without “comb” of setae; hindcoxa anteroventrally without “brush” of setae; fore
and mid trochanters without or with “brushes” of short spine-like setae; femora ven-
trally with 2 rows of thickened, usually spine-like setae; female midfemur either with
bare membranose femoral organ in its apical half, or lacking it entirely; hindtarsi in
both sexes asymmetrical, with brushes of dense setulae on medial surfaces of tar-
someres; syntergite 1+2 in both sexes narrowed at middle; female with rather short,
ventrally curved oviscape (not longer than remaining abdomen), bearing 2 sclerotized
hook-like projections ventro-apically (except in one Oriental species), but without spi-
nules or taenia-like sclerites on eversible membrane; aculeus short, flattened dorso-ven-
trally, with wide, bulky base and narrow stiletto-like apex; 3 oval, smooth spermathe-
cae; in male hypandrium narrow, with gonites flap-like microtrichose; phallapodeme
very narrow, vanes separate; ejaculatory apodeme fan-shaped; phallus without filaments
of acrophallus, with paired, sometimes loop-like sclerites of praeputium.
D i s t r i b u t i o n. Species of Eupyrgota occur in the Palaearctic, Afrotropical,
Oriental and Australasian Regions (Papuan Subregion). A preliminary study of available
material shows that there are up to 15 species of Eupyrgota in the Oriental and up to 6
in the Australasian Regions. The Palaearctic fauna includes 9 species occurring in east-
ern Asia and one, E. wagae (Bigot), in Central Asia and the Near East (see Korneyev
(2004) and Nartshuk, Korneyev (2005) for references).
T a xonomy. In the Afrotropical Region, the genus is represented by two species
groups. The first group consists of E. spinifemur (Hendel) and E. saegeri Van-
schuytbroeck, with rather slender pale yellow body and legs, moderately long (6–8 mm)
hyaline wings, a single pair of scutellar setae and small and weak hooks of the oviscape.
The second group includes E. latipennis (Walker) and 8 related species, which share
more numerous scutellar setae (two or more pairs), robust shape and reddish yellow to
brown coloration of legs, grayish or brownish microtrichose wings usually with a brown
preapical spot, body brownish red to dark brown, all setae (including ventral spines of
femora) short, and the ventral hooks of the oviscape wide and strong.
6 V. A. Korneyev
Most characters, including chaetotaxy, wing venation and morphology of female
and male terminalia, indicate that E. spinifemur is related to the Oriental and eastern
Palaearctic E. facialis Hendel. The species of the E. latipennis group are related to the
eastern Asian E. luteola, the type species of the genus, by having short setae (including
2–3 pairs of scutellars), similar venation and pattern of the wing, and the shape of the
terminalia in both sexes. Eupyrgota scioida Hendel, 1908 from the Moluccas (the type
species of Apyrgota Hendel, 1909) also belongs in Eupyrgota (Korneyev, unpublished
data), but further discussion of this nomenclatural problem, which involves several Old
World species assigned to Apyrgota, beyond the scope of the current work. Afrotropical
Apyrgota marshalli Hendel, 1914, however, is not congeneric with E. scioida and belongs
elsewhere (Korneyev, personal observation).
spinifemur group of species
Head coloration variable, from entirely yellow to mostly brown; frontal vitta matt
reddish yellow to brown with sparse, relatively long black setulae in anterior half; 1 pair
of ocellar and 1 antero-medially directed orbital setae moderately long (0.3–0.4 times as
long as eye); medial vertical seta almost as long as ocellar; frontal plate and parafacial,
including subocular plate, strongly shining, smooth, with sparse and very small, indis-
tinct setulae anterior to eye, entirely yellow, rarely with brown subocular spot or entire-
ly dark brown (fig. 1, 2, 5; 3, 1, 3); antenna yellow to brownish yellow; scape with
8–12 black antero-dorsal setulae; pedicel elongate, twice as long as wide at apex, dense-
ly setulose; first flagellomere microtrichose, narrowly elliptic in outline, sometimes
almost apex; face shining yellow, sometimes largely brown; medial carina entirely absent;
antennal grooves fused; epiclypeal sclerite very low, 0.10–0.15 times as high as remain-
ing face; frontogenal suture concolorous with face, yellow, seldom brown; parafacial
1.4–1.7 times as wide as 1st flagellomere in female, and conspicuously larger in male
(1.9–2.0 times); gena 0.2–0.25 times as high as eye vertical diameter in female and 0.4
in male; postgena and occiput yellow, rarely with brown marks along vertex and occip-
ital sutures, short microtrichose; median occipital sclerite with 0–4 postocellar setulae
and 3–4 setulae at each dorso-lateral side; 5–7 fine and short supracervical setae on each
side, half as long as other occipital setae; 30–40 occipital setae half as long as medial
vertical seta setae; one tiny lateral vertical setula indistinguishable from 7–10 setulae of
postocular row; mouth parts moderately large; palpus yellow, narrow oval, with short
blackish setulae, not dilated apically, as long as 1st flagellomere.
Thorax more or less uniformly reddish yellow or, rarely, dark brown with noto-
pleural triangle, anatergite and scutellum light yellow; postpronotal lobe with 0–2 post-
pronotal seta almost as long as vertical and ocellar setae surrounding setulae 1–7 setu-
lae ca. half as long as and; mesonotum in short and sparse microtrichia not hiding
underlying cuticle, with very sparse and short dark brown setulae; scapular setae poor-
ly differentiated; 2, occasionally 1 notopleural, 1 supra-alar, 1 intra-alar and 1 post-alar
setae present; 1 pair of scutellar setae; proepisternum with 1–5 setae as long as post-
pronotal seta and 0–5 setulae; anepisternum with 0–1 strong seta and 1–5 short black
setulae at dorsal margin of prephragmal portion; katepisternum with 0–1 seta, 0–1 dor-
solateral setulae and 1 ventral seta and 1–2 setulae at posteroventral margin;
anepimeron with 0–1 anepimeral seta and 0–1 setulae.
Legs orange-yellow, sometimes partially brown; forecoxa with 5–6 fine setulae and
1–2 stronger setae; midcoxa with 2 setae on dorso-lateral part and 1–2 seta and 2–4 se-
tulae on anteroventral margin; hindcoxa with 1 seta on dorsolateral part, and 5–7 setu-
lae; trochanters with a few (2–5) setula, foretrochanter with basal posterodorsal seta;
all femora short setulose dorsally, with one subequal basiventral seta, 1 slightly shorter
seta at basiventral one-third of femora, and with short, thickened setae ventrally, form-
ing subequal anteroventral and posteroventral rows in apical half. Midfemur of female
with or without femoral organ.
7A Revision of Afrotropical Species of the Eupyrgota (Diptera, Pyrgotidae)...
Wing hyaline, sometimes with yellowish tinge along apical half of r1 cell, often
brownish around apical portion of R2+3 vein; the latter slightly arcuate before stump
vein, then bowed anteriorly (fig. 1, 1; 3, 5–6); R4+5 bare, ending at or slightly poste-
rior of wing apex; vein M strongly bowed posteriorly. Upper calypter with 40–50 rather
thin black ciliae.
Male abdomen brownish yellow, sometimes with posterior margins of tergites yel-
low, sparsely microtrichose and brown or black setulose; 1+2nd syntergite 1.4–1.5 ti-
mes as long as tergites 3 and 4 combined, conspicuously narrowed in anterior 0.6 of its
length; 5th tergite 0.5 times as long as l+2nd syntergite.
Male postabdomen similar to that in E. tigrina Han et Kim (Kim, Han, 2005:
fig. 6) with epandrium almost as high as long in profile, and wider than high; lateral
surstylus covered with short setulae in posteroventral half; medial surstylus bar-like,
slightly curved in ventral part; ejaculator conspicuously wider than long, like in E. tig-
rina (Kim, Han, 2005: fig. 6); phallus glans with loop-like sclerite of praeputium;
proctiger dorsoventrally flattened, with short, but long setulose cerci separated by
desclerotized membrane.
Female abdomen usually uniformly yellow or with brownish areas, sparsely microtri-
chose and black setulose, with 5–7 pairs of rather long marginal setae on each tergite;
1.1–1.2 times as long as tergites 3–6 combined; sternites 4–6 each with moderately short
anteromedial apodeme (fig. 2, 1–2); oviscape (fig. 2, 1–3; 3, 3–8) shining brownish yel-
low, smooth, with short blackish setulae, 0.75–0.8 times as long as preabdominal tergites
combined; apicoventrally with a pair of weak, acute claw-like in outline, yellowish hooks,
ventrally separated, without fields of trichoid sensilla lateral and 3–4 strong setulae insert-
ed on base of hook; ventral side of oviscape long, and in postero-medial part densely
setose; aculeus (fig. 2, 6; 3, 10) basally widened, with long tapering apical part 3.5–
4 times as long as its bulky base; 3 oval spermathecae (fig. 2, 5; 3, 9).
R ema r k s. Species of this group are closely related to E. facialis Hendel and
E. tigrina Kim et Han, which share very similar wing venation, only one pair of scutel-
lar setae and shape of epandrium and phallus. The two recognized species are very si-
milar, differing by the presence or absence of the femoral organ and density of setae
and setulae. They occur sympatrically (in Nigeria and Democratic Republic of Congo)
and might represent different morphs of one polymorphic species; however, the known
material does not give any solution of this problem.
Eupyrgota saegeri (Vanschuytbroeck, 1963) comb. n. (fig. 1, 2)
Lygiohypotyphla saegeri Vanschuytbroeck, 1963: 46; Steyskal, 1972: 3; 1980: 559.
T yp e ma t e r i a l. Holotype { Lygiohypotyphla saegeri: Democratic Republic of Congo: “Congo Belge,
P.N.G. [Parc National Garamba] / Miss. H. De Saeger / II/fd/4, 3–VI–1952 / H. De Saeger, 3694 [Savane
herbeuse, I.C. 8–11 a.m., fauchages des Graminées rudérales bords de sentiers]”, “P. Vanschuytbroeck det.
/ Lygiohypotyphla / saegeri n. sp. / Holotypus [red paper label]”, “Type [pink carton label]”, (RMCA) (head
glued; antennae and 4 legs missing; abdomen dissected and pinned in microvial). Paratype }: Democratic
Republic of Congo: “Congo Belge, P. N. G. [Parc National Garamba] / Miss. H. De Saeger / PFSK. 22/8,
10–VI–1952 / H. De Saeger, 3608 [Tête de source boisée, I. A. 9–12 a. m., Fauchage de la strate d’Her-
bace´es sciaphiles]”, “P. Vanschuytbroeck det. 1962 / Lygiohypotyphla / saegeri n. sp. / Para- / type [red
paper label]”, } (De Saeger) (RBINH).
Non - t y p e ma t e r i a l. Côte-d’Ivoire: “Bouafle. Parkement / Hotel. 19.04.1989 / 06°59'N : 05°45' W /
around hotel lights / at night”, } (Londt) (NMPM); Nigeria: Zaria: Samaru, at light, 13.06.1966, };
“m.[ercury] v.[apour lamp] trap”, 31.05.1967, }; 12.06.1967, }; 24.06.1967, }; 26.06.1967, 3 }; 1.07.1967, };
10.07.1967, 2 }; 18.05.1968, }; 12.06.1968, }; 18.06.1968, }; 20.06.1968, }; 24.06.1968, }; 24.06.1968, };
26.06.1968, 3 }; 1.07.1968, }; 14.07.1968, }; 3.06.1969, }; 25.05.1972, {; 4.06.1972, }; 10.06.1972, {, };
11.06.1972, }; 16.06.1972, }; 18.06.1972, }; 19.06.1972, }; 7.07.1972, } (Deeming) (USNM).
D i a gno s i s. This species can be easily differentiated from other species of
Eupyrgota by face concave, without facial carina; female midfemur without femoral
organ. See also redescription of E. setifemur for comparison.
8 V. A. Korneyev
Rede s c r i p t i o n. Head yellow, in 5 females with brownish stripes along occipi-
tal sutures and subocular dilation; face yellow, in 2 of 3 males with pair of vertical shin-
ing black stripes; in one male frontal stripe, most of parafacial and occiput brown. Head
length : height : width ratio 1 : 1.2 : 1.3; eye ratio 0.52–0.65 in both sexes; genal–eye
height ratio 0.27–0.35 in female and 0.44–0.52 in male; flagellomere 1 almost paral-
lel-sided, narrowly or broadly rounded at apex; flagellomere 1 : pedicel ratio 1.0–1.2.
Face 1.9–2.3 times as high as wide.
Thorax of females usually yellow, sometimes with brown stripe on anepisternum
and brownish areas on anepimeron, katepisternum and katepimeron; darker in male.
Postpronotal lobe usually without seta and with a few (1–3) short setulae or, rarely with
one seta and 0–2 setulae. Mesonotum in very sparse and short setulae. Anepisternal seta
absent, at most 1–2 short setulae at postero-ventral margin of anepisternum.
Wing hyaline, cell r1 pale yellowish along costa, and brown in its broadened api-
cal portion and along apical one-quarter of vein R2+3 in female, and also with brown-
ish apical part of cell r2+3 and dark shaded apical section of vein M in some males.
Legs uniformly yellow in female and brown to brownish yellow in male. Midfemur
entirely setulose, without trace of femoral organ.
Wing length 6.5–7.5 in female and 6.0–7.2 in male.
Abdomen uniformly yellow in female (rarely syntergite 1+2 and tergites 5 and
6 slightly brownish) or brownish yellow in male. In female, synsternite 1+2 almost rect-
angular, as wide on posterior margin as on anterior, with 1–3 pairs of posterior
marginal setae and sparse setulae; sternites 2–4 transverse, twice as wide as long, with
sparse setulae and 2–4 pairs of long marginal setae; sternite 6 subrectangular, with
rounded posterior margin and 2–3(rarely to 4) pairs of long postero-marginal setae. In
male, synsternite 1+2 strongly V-shaped narrowed posteriorly, with 8–12 long and se-
9A Revision of Afrotropical Species of the Eupyrgota (Diptera, Pyrgotidae)...
Fig. 1. Eupyrgota saegeri, holotype { (RMCA) (1–3) and non-type } from Nigeria (USNM) (4–5): 1 —
habitus, right; 2 — labels; 3–5 — head (3 — right; 4 — dorsal; 5 — anterior).
Ðèñ. 1. Eupyrgota saegeri, ãîëîòèï { (RMCA) (1–3) è íåòèïîâàÿ } èç Íèãåðèè (USNM) (4–5): 1 —
îáùèé âèä, ñïðàâà; 2 — ýòèêåòêè; 3–5 — ãîëîâà (3 — ñïðàâà; 4 — äîðñàëüíî; 5 — ñïåðåäè).
veral shorter setae at posterior margin (fig. 2, 7); sternites 3 and 4 narrow, at least half
as wide as long; sternite 5 trapezoid, more than twice as wide on posterior margin as
on anterior, uniformly setulose, with two long setae at postero-marginal corner.
Male postabdomen. Epandrium as in fig. 2, 8–9; setulae on lateral surstylus sparse,
dorsalmost above proctiger; medial surstylus with 7–10 setulae. Glans of phallus as in
fig. 2, 10–11.
Female postabdomen. Oviscape reddish yellow, shining, with a few short setulae
dorsolaterally; ventral surface rather densely black setulose, with numerous long, but
comparatively thin and sparse setae in posterior half; paired hooks very thin claw like
(fig. 2, 4). Aculeus as in fig. 2, 6. Three oval spermathecae (fig. 2, 5).
10 V. A. Korneyev
Fig. 2. Eupyrgota saegeri, non-type } from Nigeria (USNM) (1–6) and holotype { (MRAC) (7–11) (from
compound microscope): 1–2 — abdomen (1 — ventral, 2 — right); 3 — abdominal sternites 3–6, dorsal,
showing anteromedial apodemes; 4 — apex of oviscape; 5 — spermathecae (2 of 3); 6 — aculeus; 7 —
abdominal sternites 1–5; 8–9 — epandrium (8 — left, 9 — posterior); 10–11 — glans (10 — right, 11 —
ventral).
Ðèñ. 2. Eupyrgota saegeri, íåòèïîâàÿ } èç Íèãåðèè (USNM) (1–6) è ãîëîòèï { (MRAC) (7–11) (ôîòî
ñî ñâåòîâîãî ìèêðîñêîïà): 1–2 — áðþøêî (1 — âåíòðàëüíî, 2 — ñïðàâà); 3 — áðþøíûå ñòåðíèòû
3–6, äîðñàëüíî, âèäíû ïåðåäíåìåäèàëüíûå àïîäåìû; 4 — âåðøèíà îñíîâíîãî ÷ëåíèêà ÿéöåêëàäà;
5 — ñïåðìàòåêè (2 èç 3); 6 — ëåçâèå ÿéöåêëàäà; 7 — 1–5-é áðþøíûå ñòåðíèòû; 8–9 — ýïàíäðèé (8 —
ñëåâà, 9 — ñçàäè); 10–11 — ãëàíñ (10 — ñïðàâà, 11 — âåíòðàëüíî).
Rema r k s. This species differs from E. spinifemur mainly by the absence of the
femoral organ in female. Other characters, e. g., the absence of anepisternal seta and
the sparser “brush” of setae on the oviscape, are somewhat variable and less definitive.
Males of E. saegeri and E. spinifemur can be distinguished only by the number of setu-
lae and setae on the thorax. The holotype male of E. saegeri is partially damaged (head
broken off and then glued to specimen, antennae and setae on head lost). The identi-
ty of this species is partially based on the female paratype, which does not have either
femoral organ, or oviscape brush and anepisternal seta. The holotype male also possess-
es no anepisternal seta. The 3 known male specimens vary in coloration from almost
entirely yellow to mainly brown with yellow pattern.
Eupyrgota spinifemur (Hendel, 1934) comb. n. (fig. 3)
Adapsilia spinifemur Hendel, 1934: 148; Steyskal, 1980: 556.
Metropina nigra Vanschuytbroeck, 1963: 25; Steyskal, 1972: 5; 1980: 559, syn. n.
T yp e ma t e r i a l. Holotype } Adapsilia spinifemur: Nigeria: “N. Nigeria / Abinsi / R. Benue /
1912–1913 Dr. J. M. Dossiel”, “spinifemur H. / det. F. Hendel <white paper>“ “Coll. Hendel” (NHMW).
Holotype { Metropina nigra: Democratic Republic of Congo: “Congo Belge: P. N. A. [Parc National Albert =
Virunga National Park] / Shamuheru (Volc. Nyamuragira) / 1843 m. 15.06.1935 / G. F. de Witte : 1434”,
“Prep. Micr. No. 2”, “P. Vanschuytbroeck det. / Metropina { / nigra n. sp. / Holotypus [red paper label]”,
“Type [pink carton label]” (RMCA).
Non - t y p e ma t e r i a l. Nigeria: Zaria: Samaru, “m.[ercury] v.[apour light] trap”, 15.07.1968, };
14.05.1972, }; 10.06.1972, } (Deeming) (USNM); same locality, “Mercury vapour / light trap”,
3–10.07.1970, }; 13–20.07.1970, } (Ward) (BMNH).
D i a gno s i s. This species, as well as E. saegeri, can be easily differentiated from
other species of Eupyrgota by concave face without facial carina; differing from
E. saegeri by midfemur with well-developed femoral organ in female, and more numer-
ous setae and setulae on postpronotum and pleuron in male (see key to species above).
Red e s c r i p t i o n. Similar to E. saegeri, differing as follows. Head yellow, in 2 fe-
males with pale brown stripes on occiput; in male with entirely brown frontal vitta and
partially brown face, parafacial (including subocular dilation) and occiput. head length
: height : width ratio 1 : 1.2–1.3 : 1.2–1.3 (in most specimens somewhat wider than
high, in some slightly higher than wide); eye ratio 0.61–0.64 in female and 0.65 in
male; genal–eye height ratio 0.29–0.34 in female and 0,35 in male; flagellomere
1 slightly narrowed to apex or almost parallel-sided and bluntly rounded; flagellome-
re 1 : pedicel ratio 1.0–1.1. Face 1.8–2.3 times as high as wide.
Thorax of females yellow, with brown stripe on anepisternum; in male more
brownish (cuticle hidden). Postpronotal lobe 1 seta and 2–7 short setulae. Mesonotum
in sparse and moderately short setulae. Anepisternal seta present, but no short setulae
on postphragmal part in all studied specimens (2–8 setulae at dorsal margin present).
Wing hyaline, with cell r1 along costa and vein R2+3 yellow, rarely darker brown
apex of r1 cell, in female; in male also with yellow bordered apical sections of veins
dm-cu, R2+3 and M.
Legs yellow in female and brownish yellow in male. Midfemur in its distal third
quarter with femoral organ 0.25 times as long as femur.
Wing length 6.8–7.5 in female and 7.9 in male.
Female abdominal synsternite 1+2 slightly narrower on posterior margin than on
anterior; sternite 6 with 3–4 pairs of long postero-marginal setae. In male, sternite
5 with 4–5 pairs of long setae. Male postabdomen not examined.
Female postabdomen. Oviscape on ventral surface with numerous thick and dense
setae in posterior half (fig. 3, 7–8); paired hooks slightly flattened, sickle-like (fig. 3,
7–8). Aculeus as in fig. 3, 10. Three oval spermathecae (fig. 3, 9).
Commen t s. The holotype of M. nigra is entirely covered with mud of unknown
origin, which hides the coloration of the cuticle and wing pattern, which misled
11A Revision of Afrotropical Species of the Eupyrgota (Diptera, Pyrgotidae)...
Fig. 3. Eupyrgota spinifemur, non-type } from Nigeria (USNM) (1–5, 7–10) and Metropina nigra, holotype
} (MRAC) (6): 1 — head and anterior portion of thorax, left; 2 — head and mesonotum, dorsal; 3 — head,
anterior; 4 — midfemur, anterior; 5–6 — wing; 7 — abdomen, ventral; 8 — oviscape; 9 — spermathecae;
10 — aculeus.
Ðèñ. 3. Eupyrgota spinifemur, íåòèïîâàÿ } èç Íèãåðèè (USNM) (1–5, 7–10) è Metropina nigra, ãîëîòèï
} (MRAC) (6): 1 — ãîëîâà è ïåðåäíÿÿ ÷àñòü ãðóäè, ñëåâà; 2 — ãîëîâà è ñðåäíåñïèíêà, äîðñàëüíî; 3 —
ãîëîâà, ñïåðåäè; 4 — ñðåäíåå áåäðî, ñïåðåäè; 5–6 — êðûëî; 7 — áðþøêî, âåíòðàëüíî; 8 — îñíîâíîé
÷ëåíèê ÿéöåêëàäà; 9 — ñïåðìàòåêè; 10 — ëåçâèå ÿéöåêëàäà.
P. Vanschuytbroeck to conclude that it is entirely dark colored (including wings). I
have cleaned several parts on the preliminarily softened holotype and observed that it
is brown to pale brown with yellow pattern similar to male specimens of E. saegeri. It
has 5–7 setulae on the postpronotal lobe and well-developed anepisternal setae on both
sides, which indicates that this specimen is conspecific with the female holotype of
Adapsilia spinifemur rather than with E. saegeri.
latipennis group of species
Des c r i p t i o n. Head yellow to partially brown; frontal vitta matt reddish yellow
to brown with sparse setulae; anterior part with longer anteriorly directed setulae above
lunule; 1–3 pairs of ocellar and 1 antero-medially directed orbital setae; medial verti-
cal seta as long as ocellar; frontal plate and parafacial matt, faintly shagreened to waxy
shining, shallowly pitted or wrinkled, rarely completely smooth, with numerous tiny
12 V. A. Korneyev
setulae almost to ventral margin of eye (fig. 4, 2, 4; 7, 7–9; 9, 3–4); antenna yellow
to dark brown; pedicel densely setulose; first flagellomere microtrichose, narrowly ellip-
tic in outline, usually slightly tapering to apex; face yellow, usually with dark brown or
black areas; medial carina well developed, narrow, dilated ventrally, matt, 0.7–0.85 ti-
mes as long as facial height; antennal groove shining; epiclypeal sclerite low,
0.20–0.45 times as high as facial ridge, matt or subshining, often with transverse wrin-
kles (fig. 7, 9; 9, 3–4); frontogenal suture narrowly black in ventral part; parafacial
1.5–3.2 times as wide as 1st flagellomere; gena 0.28–0.53 as high as eye vertical dia-
meter, subocular dilation subshining or shining, often with dark subocular spot; post-
gena and occiput yellow to brown, moderately densely microtrichose; median occipital
sclerite with 1–4 pairs of postocellar setulae and 3–6 setulae at each dorso-lateral side,
often with one or two oblique brown marks in the middle, forming triangular, Y- or V-
shaped mark; up to 20–30 fine and moderately long supracervical setae on each side,
almost as long as other major head setae; 35–50 occipital setae 1.5 times as long as
postocular setae; up to 20–25 setae in each postocular row; mouth parts very large; pal-
pus yellow with short blackish setulae, apically dilated and bluntly rounded, as long as
1st flagellomere.
Thorax usually pale yellow, often with reddish yellow mesonotal scutum and post-
notum, and brownish band on anepisternum and katepimeron; in darker specimens most-
ly brown, with yellow postpronotal lobe, scutellum, posterior part of anepisternum, most
of katepisternum, and whole katatergite; postpronotal lobe with 10–20 setulae ca. half as
long as vertical and ocellar setae and 0–2 postpronotal seta almost twice as long as sur-
rounding setulae; mesonotum in short and sparse microtrichia not hiding underlying cuti-
cle, with sparse dark yellow or brown setulae; scapular setae poorly or not differentiated;
2 notopleural, 1 supra-alar, 1 intra-alar and 1 post-alar setae present; 2–3 pairs of scutel-
lar setae (rarely up to 5 pairs); proepisternum with 8–22 setulae; anepisternum with
1 strong seta and 6–7 long black setulae along posterior margin and 5-10 setulae at dor-
sal margin of prephragmal portion; katepisternum with 1 seta, 3–10 dorsolateral setulae
half as long as katepisternal seta and 2 stout ventral setae and 3–8 setulae at posteroven-
tral margin; anepimeron with 1 anepimeral seta and 3–10 long setulae.
Legs reddish yellow to reddish brown, sometimes partially dark brown; forecoxa
with 12–15 fine setulae and 1–2 stronger setae; midcoxa with 1 strong seta on dorso-
lateral part and row of 6–7 strong unmodified setulae 0.9–1.0 times as long as longest
seta on anteroventral margin; hindcoxa with 1 strong seta on dorsolateral part, and
20–40 strong, posteroventrally directed setae on anteroventral margin 0.7–0.9 times as
long as seta, forming neither comb, no brush; trochanters with fine setulae in male; in
female, fore- and midtrochanters sometimes with short spine-like setulae forming
brush; hindtrochanter fine and sparsely setulose; all femora with subequal 1 basiventral
seta, 1 slightly shorter seta at basiventral one-third of femora and with thickened, but
moderately long setulae ventrally, forming subequal anteroventral and posteroventral
rows in apical half; forefemur sometimes with strong dorsal setae.
Wing hyaline, sometimes with yellowish tinge in anterior half or widely brownish;
antero-apical part with brown spot (fig. 9, 6) or widely brown in anterior half; R4+5
bare or, very rarely, setulose to r-m crossvein. Upper calypter with 7–8 thickened cil-
iae at postero-distal margin.
Male abdomen reddish yellow to dark reddish brown, sparsely microtrichose and
brown or black setulose; 1+2nd syntergite 1.1–1.5 times as long as tergites 3 and
4 combined, often with light yellow band, or “clitellum” at middle; 5th tergite
0.5–0.8 times as long as l+2nd syntergite. Male terminalia (fig. 6, 8, 6–10; 9, 13–15)
similar to those in E. wagae (Bigot) (Korneyev, 2004: fig. 16) with epandrium higher
than long in profile; lateral surstylus densely covered with thick but short setulae; medi-
al surstylus bar-like, free in ventral half, with 5–10 short setulae; hypandrium narrow,
with gonites microtrichose; phallapodeme very narrow, vanes separate; ejaculatory
13A Revision of Afrotropical Species of the Eupyrgota (Diptera, Pyrgotidae)...
apodeme fan-shaped; phallus glans with asymmetrical sclerites of acrophallus; proctiger
dorsoventrally flattened, with cerci separated by desclerotized area, but not lobate.
Female abdomen (fig. 5, 1; 7, 1) of reddish yellow to brown ground color, sparse-
ly microtrichose and reddish brown or black setulose; syntergite 1+2 usually with yel-
low crossband, 0.9–1.7 times as long as tergites 3–6 combined; tergites 3–6 uniformly
brownish red to brownish yellow; oviscape subshining reddish yellow to brown, often
wrinkled, with short pale brown to reddish yellow setulae, 0.6–0.75 times as long as
preabdominal tergites combined; apicoventrally with a pair of strong, apically flattened
and often rounded in outline, black hooks, joined by V-shaped apodeme, with field of
trichoid sensilla lateral to hook and group of 6–10 setulae anterior to hook; aculeus
(fig. 5, 7; 8, 5; 9, 12) dorso-ventrally flattened, basally widened, tapering toward apex;
spermathecae (fig. 5, 3; 8, 4; 9, 11) yellowish brown, oval or sausage-like.
latipennis subgroup
Dia gno s i s. Parafacial matt, finely shagreened, usually with distinct and numerous
small pits (fig. 4, 2; 7, 9). Supraclypeal sclerite conspicuously wrinkled, high, 1.1–1.2 times
as high as width of antennal groove. Wing with 2 distinct brown spots (on pterostigma and
on antero-distal part), separated by pale yellowish area distal of R1 apex (fig. 9, 6).
S p e c i e s i n c l u d ed. E. caffra, E. latipennis and E. sublatipennis.
Eupyrgota caffra (Hendel, 1914) (fig. 4–6)
Adapsilia caffra Hendel, 1914: 90; 1934: 149; Steyskal, 1980: 556; Eupyrgota caffra: Enderlein, 1942: 120 (pro
parte; dubious identification); Adapsilia rugosigenis Hendel, 1934: 149, syn. n.; Adapsilia (Eupyrgota) rugosi-
genis: Steyskal, 1980: 557; Eupyrgota rugosigenis: Enderlein, 1942: 120 (pro parte; dubious identification);
Adapsilia vespiformis: Vanschuytbroeck, 1963: 61 (pro parte; misidentification).
Non Adapsilia caffra sensu Vanschuytbroeck, 1963: 64 (misidentification of E. latipennis).
T yp e ma t e r i a l. Holotype } Adapsilia caffra (double-mounted on celluloid rectangle, left pedicel and
1st flagellomere missing; right foreleg glued): South Africa: “Port Natal [= Durban] / Guenzins / 60–97”,
“Pt / Natal [paper circle]”, “Campylocera / caffra H. / det. Hendel”, “} Type [red-boarded circle]”,
“Holotype [red-boarded circle]”, “Holotype / Adapsilia / caffra / Hendel / verified by J. E. Chainey, 2002”,
“BMNH(E) # / 252160” (BMNH). Syntype } Adapsilia rugosigenis: Malawi: “Mlanje [= Mulanje?]”, “coll. Hen-
del”, 17.02.1913, (Neave) (“Adapsilia / rugosigenis / H. } / Hendel det.”) (NHMW);
Non - t y p e ma t e r i a l. Botswana: “Zibadianja / Noamiland / Botswana”, 24.10.1991, } (Wooff)
(BMNH); Democratic Republic of Congo: “Terr. Yahuma, 810 m”, 12.1948, } (Benoit) (RMCA) (“P. Van-
schuytbroeck det 1963 / Peltodasia / vespiformis End.”); “N. Lac Kivu: Rwankwi”, 05.1948, { (“P. Van-
schuytbroeck det 1963 / Peltodasia / vespiformis End.”) (Leroy) (RMCA); Malawi: “Nyassaland / Cholo”,
{ (Wood) (DEI), same labels, 2700 ft., {, } (CMNH); same labels, 30.10.1928, } (without femoral organ)
(Wood) (CMNH); “SE 222 GBD / Farmers Brigade 5 kms / SE of Serowe Hillside / N slope / Mercury
vapour lamp”, 28.11.1990, { (Forchhammer) (NMPM); Namibia: “Bwabwata Park / Bum Hill Campside
(Kwando R.) / 17°46'52'' S 23°20'28'' E / Malaise trap”, 10–13.02.2004, } (Kirk-Spriggs) (SMWN);
“Kavango, Popa Falls im / Kavango: 18°07'S/21°35'E”, 19–22.01.1999, }; 11.03.1992, } (Koch) (ZMHB);
Nigeria: “73 at light / Univ. coll. / Ibadan, J. R.”, 3.06.1955, } (Malloch ?) (USNM); South Africa:
Northern Province: “Nwanedi Resort, Venda, 22°38'S 30°24'E; / 550 m / R. Oberprieler / collected / at
light”, 5–9.02.1994, } (Oberprieler); “Hans Merensky Nat. / Res. 23°40' S 30°39' E”, 27–30.11.1981, }
(Mansell); “Zoutpansberg / Wyllies Poort”, 26.01.1954, { (Munro); “Ben Alberts Nat. Res. / Thabazimbi /
24°27'S 27°23'E”, 24–28.11.1980, { (Mansell) (SANC); “Duiwelskop [Duiwelskloof]”, 24.11.1948, } (Omer
Cooper) (BMNH); “Letaba camp / K. N. P. Survey”, 14–18.11.1961, } (Vari et Rorke) (NMPM); “Fiesta
Motel 20 km S[W] / Potgietersrus 24°17' S / 28°51' E; 1100 m”, “collected / at light”, 16.11.1992, 4 }
(Ho�lzel, Ohm et Mansell); same locality, 26.02.1995, 2 { (Ho�lzel, Ohm et Mansell) (SANC); Mpumalanga:
“Skukuza / Transvaal / 23.11.1959”, 2 }; “De Hoop 203 JU / Nelspruit Distr.”, 20–21.11.1972, } (Strydom)
(NMPM); Montrose Falls, 25°25'S 30°44'E, collected at light, 29.01.1989, } (Oberprieler) (SANC);
“Nelspruit”, 12.1914, } (Breijer) (RNHL); Gauteng: Pretoria, 6.02.1919, } (Munro), same locality,
1.12.1937, 2 } (van Son) same locality, 31.01.1955, } (Stuckenberg) (NMPM); same locality, Roodeplaat,
bait trap, 01.1958, 4 {, 3 }; same locality (“Roodeplaat / Pretoria Distr.”), 3.11.1958, {, }, 12.1958, { (Bot);
same locality (“Ze: Rp 237 / Loc: HRI Pretoria”), 01.1975, {, } (Donaldson) (SANC); North-West
Province: “Rustenburg Nature / Res. 25°40'S 27°12'E”, 23–26.02.1981, { (Miller) (SANC); Mpumalanga:
“Barberton”, 12.1978, { (Eardley) (SANC); KwaZulu-Natal: “Lake Sibaya, Baya / Camp 27°23'S / 32°41'E;
50 m”, collected at light, 30.11–2.12.1992, } (Ho �lzel, Ohm et Mansell) (SANC); “Percy Fife Nat. Res. /
Potgietersrus Distr.”, 18–20.01.1971, { (Jones); “Mtunzini / Zululand”, } (Schofield); “Mkuzi Game
14 V. A. Korneyev
Reserve / ca 140 m / 27°38'20'' S : 32°09'30'' E / MV light et Malaise”, 8–15.10.1990, { (Londt); “Stellab.
[Stellabush near Durban] / 17–16 / Woelig [??? - unreadible]”, } (NMPM); “Hluhluwe Game Res. / 28°02'S
: 32°05' E”, 4–6.02.1994, { (Schumann) (ZMHB); “Richards Bay”, “at night into vehicle”, 3.02.1963, 2 {
(Tinley) (NMPM); Tanzania: [D. O.-Afrika, Bumbuli (Ost Usambara) — according to Enderlein (1942):
blue label with ink faded or washed away], 5.12.1890, { (Meinhof) (ZMHB); “N.-Nyassa / Langenburg”,
03.1898, { (Fu �llerborn) (ZMHB) (“Eupyrgota rugosigenis (Hend. 1934) { / Dr. Enderlein det. 1940”);
Zimbabwe: “Yumbu Mts.”, 03.1930, } (USNM) (“Adapsilia / ypsilon / Hend. / Det. / J. R. Malloch”);
“Basely Br., S. R.”, 28.01.1963, 2 } (Cookson) “Zimbabwe / S. Rhodesia, B. R. S. P. G.”, 28.01.1955, 2 }
(NMPM); “S. Rhodesia / Vleiplaats Est. / 1952 / Rhod. Whattle bo. / prob. par. of. / Eulebida mashona”,
“Com. Inst. Entomol. / Coll. No 12955”, “Pres. by / Com. Inst. Ent. / B. M. 1953-354 [locality not found;
actually SAR?]”, { (“Adapsilia (?) sp. indet. (teneral) van Emden det. 1953”) (BMNH).
D i a gno s i s. Parafacial conspicuously pitted and wrinkled; thorax brownish yellow
to brownish red; female fore- and midtrochanter sparsely spinulose, forefemur thick-
ened, with strong dorsal setae, midfemur with femoral organ; oviscape with thin, long
and yellowish setulae anterior to hooks; aculeus with wide basal (twice as wide as base
of apical part) and long apical part (at least 3 times as long as basal); male: phallus
glans with almost symmetrical loop-like sclerites of praeputium surrounding gonopore.
Red e s c r i p t i o n. Body brownish yellow to brown without contrasting brown pat-
terns; setae and setulae brown to black; mesonotum length 2.2–4.8 mm (holotype
4.33 mm), wing length 8.3–14.5 (holotype 12.8 mm).
Head brownish yellow to light brown with frontal-head ratio 0.43–0.55, eye ratio
0.45–0.50, genal-eye ratio 0.33–0.4 in female, 0.38–0.50 in male; first flagellomere-
pedicel ratio 0.85–1.33; medial vertical seta 0.10–0.16 times as long as longest diame-
ter of eye; lateral vertical seta 0.3–0.4 times as long as medial vertical seta, often indis-
tinguishable; ocellar seta 0.8– 0.9 times as long as medial vertical seta, often lacking;
orbital seta 0.7–0.9 times as long as medial vertical seta; frons whitish to brownish yel-
low; frontal vitta matt with sparse setulae and dark brown dots at their bases, in holo-
type with large brown patch in middle; frontal plate and parafacial matt (shagreened)
or, rarely, subshining, shallowly pitted and sometimes wrinkled, with small black spot
and short brown setula in bottom of every pit; antenna yellowish brown to brown; first
flagellomere brown, densely microtrichose, 2.8–3.2 times as long as wide, narrowly ellip-
tic in outline, slightly tapering to apex; arista bare; face shining brownish yellow with
black spots between bases of antennae and two black strikes along medial side of anten-
nal groove; frontogenal suture narrowly black in ventral part; median carina 0,7 times as
long as facial height; parafacial subshining, pitted and setulose as described above,
1.4–1.7 times as wide as 1st flagellomere in male, 1.51.6 times in female; gena with matt,
light brown genal groove and subshining yellowish brown dorsal half and dark brown
subocular spot; genal dilation linear, black; postgena brownish yellow; occiput with
18–20 yellowish supracervical setulae on each side, often with two oblique brown marks
in the middle, forming incomplete V-shaped mark, brown along upper occipital suture,
and pair of brown supracervical spots or without them, or entirely yellow; mouth parts
brown, large, as long as antenna, moderately long setulose; palpus as long as flagello-
mere 1, reddish yellow with short blackish setulae, very slightly widened before apex.
Thorax reddish yellow to yellowish brown; postpronotal lobe yellow with
17–20 brown setulae approximately as long as vertical and ocellar setae and 1, rarely
2 postpronotal seta(-e) almost twice as long as surrounding setulae; scutum reddish yel-
low with short black or brown setae and moderately short and dense brownish red or
yellow setulae; scutellum reddish to brownish yellow; 2–3 pairs of scutellar setae; pleu-
ra reddish to brownish yellow ground color; presternum with cluster of 3–5 black or
brown setulae on apex of each lateroventral process; proepisternum with 12–20 mo-
derately long setulae, 3–4 dorsal longest; anepisternum of reddish yellow ground color
darker patches, with 1 seta (broken off on both sides in holotype) and 14–17 moder-
ately long setulae; katepisternum with 1 seta, 10–12 moderately long dorsolateral setu-
15A Revision of Afrotropical Species of the Eupyrgota (Diptera, Pyrgotidae)...
lae half as long as katepisternal seta and 3–4 rather stout ventral setae (partially dam-
aged on both sides in holotype) and 3–4 finer setulae at posteroventral margin;
anepimeron with 1 anepimeral seta and 7–8 long setulae; katepimeron, meron, katater-
gite and anatergite, sub-scutellum and mediotergite subshining, sparsely and short
microtrichose, pale to dark reddish yellow, with more or less distinctive wrinkles.
Legs uniformly reddish yellow; forecoxa with 15–20 fine setulae and 2–3 stronger
setae; midcoxa with 1 strong seta on dorso-lateral part, 7–8 fine and short setulae on
postero-ventral margin, and cluster of 6–8 moderately strong unmodified setulae
16 V. A. Korneyev
Fig. 4. Eupyrgota caffra, holotype } (BMNH) (1–3), syntype } A. rugosigenis (NHMW) (4–5) and non-type
} from Natal (NMPM) (6–13): 1 — habitus, left; 2, 4 — head (2 — left, 4 — dorsal); 3, 5 — labels; 6 —
coxae and trochanters, ventral; 7 — foretrochanter, anteroventral; 8 — midtrochanter, anterior; 9 —
foretrochanter and forefemur, posterior; 10–13 — midfemur, anterior (10 — total view, with coxa, trochanter
and part of tibia, 11 — femoral organ, 12 — area dorsal to femoral organ, enlarged, 13 — spurious seta of
ventral row, enlarged) (1–6 — from dissecting microscope, 7–13 — SEM).
Ðèñ. 4. Eupyrgota caffra, ãîëîòèï } (BMNH) (1–3), ñèíòèï } A. rugosigenis (NHMW) (4–5) è íåòèïî-
âàÿ } èç Íàòàëÿ (NMPM) (6–13): 1 — îáùèé âèä, ñëåâà; 2, 4 — ãîëîâà (2 — ñëåâà, 4 — äîðñàëüíî);
3, 5 — ýòèêåòêè; 6 — òàçèêè è âåðòëóãè, âåíòðàëüíî; 7 — ïåðåäíèé âåðòëóã, ïåðåäíåâåíòðàëüíî; 8 —
ñðåäíèé âåðòëóã, ñïåðåäè; 9 — ïåðåäíèé âåðòëóã è ïåðåäíåå áåäðî, ñçàäè; 10–13 — ñðåäíåå áåäðî,
ñïåðåäè (10 — îáùèé âèä, ñ òàçèêîì, âåðòëóãîì è ÷àñòüþ ãîëåíè, 11 — ôåìîðàëüíûé îðãàí, 12 —
ó÷àñòîê êâåðõó îò ôåìîðàëüíîãî îðãàíà, óâåëè÷åíî, 13 — øèïèê âåíòðàëüíîãî ðÿäà, óâåëè÷åíî)
(1–6 — ôîòî ñ áèíîêóëÿðà, 7–13 — ôîòî ñî ñêàíèðóþùåãî ýëåêòðîííîãî ìèêðîñêîïà).
0.5–0.7 times as long as longest seta on anteroventral margin, forming neither comb
nor brush; hindcoxa with 1 strong seta on dorsolateral part, and 10–15 moderately
strong setulae 0.6–0.7 times as long as seta, on anteroventral margin; in female: fore-
trochanter with moderately dense brush of thickened and short, spine-like, posteriorly
directed setulae; midtrochanter with similar, but slightly larger brush of posteriorly
directed setulae; hindtrochanter fine and sparsely setulose; in male, all trochanters ven-
trally with fine sparse setulae; all femora with slender basiventral seta, longest on forefe-
mur (as long as width of femur), with clusters of dorsoapical setae 1.5 as long as short
dorsal setulae, and with short spine-like setulae ventrally, forming subequal anteroven-
tral and posteroventral rows in apical half; forefemur conspicuously thickened in
17A Revision of Afrotropical Species of the Eupyrgota (Diptera, Pyrgotidae)...
Fig. 5. Eupyrgota caffra, non-type } from Natal (NMPM): 1 — abdomen, ventral; 2 — sternites 2–6; 3 —
spermathecae (2 of 3); 4 — apex of oviscape, eversible membrane and exposed aculeus, right; 5–6 — apex
of oviscape (membrane inverted); 7 — aculeus (1, 6 — SEM; 4 — from dissecting microscope; 2, 3, 5, 7 —
from compound microscope).
Ðèñ. 5. Eupyrgota caffra, íåòèïîâàÿ } èç Íàòàëÿ (NMPM): 1 — áðþøêî, âåíòðàëüíî; 2 — ñòåðíèòû
2–6; 3 — ñïåðìàòåêè (2 èç 3); 4 — âåðøèíà îñíîâíîãî ÷ëåíèêà ÿéöåêëàäà, âûâåðíóòûå íàðóæó
ìåìáðàíà è ëåçâèå ÿéöåêëàäà, ñïðàâà; 5–6 — âåðøèíà îñíîâíîãî ÷ëåíèêà ÿéöåêëàäà (ìåìáðàíà
ââåðíóòà âíóòðü); 7 — ëåçâèå ÿéöåêëàäà (1, 6 — ôîòî ñî ñêàíèðóþùåãî ýëåêòðîííîãî ìèêðîñêîïà;
4 — ôîòî ñ áèíîêóëÿðà; 2, 3, 5, 7 — ôîòî ñî ñâåòîâîãî ìèêðîñêîïà).
female, 3.0–3.3 times as long as wide, dorsally with 7–8 semi-appressed setae 1.5 times
as long as surrounding setulae and with 2–4 slender setae posteroventrally in basal half
next to basiventral seta (fig. 4, 9); midfemur with femoral organ half as long and almost
half as wide as midfemur in female; 3 rows of short spinulose setulae dorsal to femoral
organ (fig. 4, 10–12); very rarely, femoral organ reduced or absent; in male, forefemur
not conspicuously thickened, only 3.8–4.2 times as long as wide.
Wing hyaline with grayish microtrichiae in posterior half, with yellowish tinge in
anterior half (basal costal, costal, subcostal, basal radial and basal halves of r1, r2+3 and
antero-basal quarter of r4+5 cells); apices of r1, r2+3 and antero-apical portion of r4+5
cells with brown subapical spot; wing-thorax ratio 3.0–3.2, vein R4+5 ratio 1.5–1.8,
vein M ratio 0.40–0.68.
Male abdomen reddish yellow to reddish brown, sparsely microtrichose and brown,
short setulose; syntergite 1+2 1.2–1.3 times as long as tergites 3 and 4 combined; and
1.8–2.0 times as long as tergite 5; synsternite 1+2 long and narrowed to posterior mar-
gin, 5.0–5.2 times as long as wide at posterior margin; epandrium and hypandrium as
in fig. 6, 1; glans of phallus with almost symmetrical sclerites of praeputium of loop-
like appearance; lateral flap of glans corpus short and almost appressed (fig. 6, 2–3).
Female abdomen similarly colored; syntergite 1+2 1.2–1.4 times as long as tergites
3–6 combined; oviscape subshining yellowish or reddish brown ground color, robust,
0.74–0.78 times as long as preabdominal tergites combined and slightly wider than long
on ventral side; apicoventrally with pair of rather soft lobes, densely covered by trichoid
sensillae and closing oviscape aperture in rest; pair of strong black hooks medial to each
inverted in rest and exposed ventrobasally of everted membrane; ventral surface of ovis-
cape anterior to hooks with 6–8 short fine yellow setulae; aculeus with base gradually
transiting into long stiletto-like piercing part more than 2.9–3.4 times as long as its
bulky base (fig. 5, 7); spermathecae elongate oval to sausage-like, 3.5–7 times as long
as wide (fig. 5, 3), one of 3 sometimes V- or Y-like bifurcated.
Rema r k s. E. caffra is widespread mainly in southern Africa, but a single
undoubtedly identified female is recorded from Nigeria. The femoral organ, a structure,
which differentiates E. caffra from E. latipennis and E. sublatipennis is absent in one
female from Malawi (which fits the diagnosis of E. caffra in genitalic characters).
F. Hendel (1934) gave only a brief diagnosis of A. rugosigenis in the key: “[Fronto-
orbital plates] in the anterior portion of the frons with transverse wrinkles. Parafacial
twice as wide as flagellomere 1; antennal groove only 2/3 as long as face… 11–16 mm.
Uganda” (translated from German). The only specimen in Hendel’s collection
(NHMW) under this name is a large female of A. caffra from Mlanje (Malawi) (see
18 V. A. Korneyev
Fig. 6. Eupyrgota caffra, non-type { from Natal (NMPM): 1 — male terminalia, right (segments 6–8 and
phallus detached); 2–3 — glans (2 — right, 3 — ventral).
Ðèñ. 6. Eupyrgota caffra, íåòèïîâîé { èç Íàòàëÿ (NMPM): 1 — òåðìèíàëèè ñàìöà, ñïðàâà (6–8-é
ñåãìåíòû è ôàëëþñ óäàëåíû); 2–3 — ãëàíñ (2 — ñïðàâà, 3 — âåíòðàëüíî).
above). It therefore did not originate from “Uganda”, and the original statement of
Hendel is a lapsus calami (like in several other places in the same paper); this speci-
men is obviously a possibly unique syntype of E. rugosigenis.
Eupyrgota latipennis (Walker, 1849) (fig. 7, 8)
Oxycephala latipennis Walker, 1849: 1087; Adapsilia latipennis: Hendel, 1914: 88; 1934: 148; Vanschuytbroeck,
1963: 65; Adapsilia (Eupyrgota) latipennis: Steyskal, El-Bialy, 1968: 39; Steyskal, 1972: 1; Steyskal, 1980: 557.
Adapsilia ypsilon Hendel, 1914: 89; 1934: 149; Steyskal, 1972: 1; Adapsilia (Eupyrgota) ypsilon: Steyskal, 1980:
557; Eupyrgota ypsilon: Enderlein, 1942: 119, syn. n.
Adapsilia rugosigenis Hendel, 1934: 149, possible synonym. Eupyrgota rugosigenis: Enderlein, 1942: 119 (mis-
identification).
Adapsilia vespiformis: Vanschuytbroeck, 1963: 61 (pro parte; misidentification).
Non Adapsilia ypsilon: Vanschuytbroeck, 1963: 64 (misidentification of Clemaxia sp. near gracilis).
T yp e ma t e r i a l. Holotype } Oxycephala latipennis: Sierra Leone: “Sierra Leone”, “38 / 11. 8 / 295
[circle]”, “Type [green bordered circle]”, “Holo / type [red bordered circle]”, “one of Walker series so
named”, “Holotype Oxycephala latipennis Walker verified J. E. Chainey 2002”, “BMNH # 25167”. Double-
mounted on celluloid rectangle, right pedicel and 1st flagellomere missing; apex of right wing partially broken
off. (BMNH). Holotype } Adapsilia ypsilon: Sierra Leone: “130 / Sierra Leone / Protectorate, / West Africa.
/ March 1910 / Dr. H. E. Arbuckle. / 1910-247”, “} / Type [red bordered circle]”, “Eupyrgota / ypsilon,
H. [Hendel’s handwriting] / det. Hendel”, “Holotype [red bordered circle]”, “Holotype Adapsilia ypsilon
Hendel Verified by J. E. Chainey, 2002”, “BMNH(E) # 252166” (BMNH).
Non - t y p e ma t e r i a l. Burundi: “Urundi: Rugari, Dames de Marie”, 1948, } (“P. Vanschuytbroeck
det 1963 / Peltodasia / vespiformis End.”) (RMCA); “Urundi: Kisenyi”, “R. I. Sc. N. B. / I. G. 24.452”,
05.1956, } (Franc �ois) (RBINH); Côte-d’Ivoire: “Javier / Adiopodoume”, “50”, 7.09.1949, { (collector
unknown); “Dimbroko”, 1919, } (oviscape lacking) (Seguy) (MNHNP); Democratic Republic of Congo:
“Kibali-Ituri: Bayenga, Terr. Wamba, 810 m”, 01.1956, } (“P. Vanschuytbroeck det 1963 / Adapsilia / caffra
Hend.”) (RMCA); Rutshuru, “R. Mus. Hist. Nat. /. Belg. I. G. 10.482”, 04.1937, } (Ghesquiere) (IRSNB);
Kibali-Ituri: Mahagi-Niarembe, 09–10.1935, 3 { (“P. Vanschuytbroeck det 1963 / Peltodasia / vespiformis
End.”) (Scops) (RMCA); “Rutshuru, 1285 m : 118”, 25.11–20.12.1933, { (“P. Vanschuytbroeck det 1963
/ Peltodasia / vespiformis End.”) (de Witte) (RBINH); Kivu: Buseregenye (Rutshuru), 09.1929, { (“P.
Vanschuytbroeck det 1963 / Peltodasia / vespiformis End.”) (Luja) (RMCA); Kivu: Lwiro, 26.11.1966, },
11–12.1966, 3 } (“Peltodasia latipennis (Walk.) d. Steyskal 69”) (Jilly) (SMNS); “Kapiri / Miss. Agric.”,
09.1912, }; Lulua: Kapanga, 05–06.1959, } (Allaer) (“P. Vanschuytbroeck det 1963 / Peltodasia /
vespiformis End.”) (RMCA); Ethiopia: “Abessinien / Dire Daoua” [= Dire Dawa], 3.06.1936, { (“Eupyrgota
/ rugosigenis / (Hend. 1934) { / Dr. Enderlein det. 1940”) (Uhlenhuth) (ZMHB); Gambia: “Keneba, at
mercury light”, 19.06.1974, 2 {, } (BMNH); Ghana: “Gold Coast: / nr. Accra”, 9.07.1943, { (Guichard)
(BMNH); Malawi: “Zomba Plateau / 1500 m, 1535 Ad Montane forest”, 12–14.12.1980, } (Londt et
Stuckenberg) (NMPM); Namibia: “Kavango: Pope Falls im / Kavango [Kaprivi Strip], 18°07' S / 21°35' E”,
13.03.1992, } (Koch) (ZMHB); Tsumkwe District: “/ Trekkersboom / 19°18'00'' S 20°39'42'' E / sweeping
grasses in wet area”, 26.11.1998, } (Kirk-Spriggs); Tsumeb District: “Varianto 771/2 / 19°23'00'' S 17°43'57''
E / light trap”, 28.03–01.04.2001, } (Kirk-Spriggs et Mey); Otjinene District: “Epukiro River, 3 km N at:
/ 21°22'26'' S 20°06'09'' E / Malaise trap samples”, 09–11.02.2001, } (Kirk-Spriggs, Marais et Wheeler); “De
Hoek 838 / 21°56'26'' S 20°58'55'' E / Malaise trap samples”, 03–06.02.2001, } (Kirk-Spriggs, Marais et
Wheeler) ; Gobabis District: “Somerkoms 521 / 22°01'59'' S 19°57’22'' E / Malaise trap samples”,
06–08.02.2001, 3 } (Kirk-Spriggs, Marais et Wheeler); “Kaudom Game Pk. / 10 km W of Dussi / 18°48'32'' S
20°43'57'' E / Malaise traps”, 29–30.12.1998, } (Kirk-Spriggs, Marais et Mann) (SMWN); Nigeria: Samaru,
“Mercury vapour light trap”, 3–10.06.1970, }; 15–22.06.1970, }; 13–20.07.1970, }; 21– 29.07.1970, }
(Ward) (BMNH); “Zaria, Samaru”, “m. v. trap”, 26.05.1967, }, 19.06.1972, { (dissected) (Deeming)
(USNM); “73 at light / univ. coll. / Ibadan / J. R.”, 3.06.1955, } (Malloch?) (USNM); Rwanda: “Kisenyi
1500 m”, 20.01.1952, } (“P. Vanschuytbroeck det 1963 / Adapsilia / latipennis / Walk.”); “Kisenyi”,
10.1952, } (“P. Vanschuytbroeck det 195... / Adapsilia / latipennis Walk.”); “Kigali / 1500–1800 m”,
12.10.1951, } (“P. Vanschuytbroeck det 1963 / Adapsilia / latipennis / Walk.”) (Bertrand) (RMCA); Saudi
Arabia: Melalu, 2.11.1948, } (Watson); “Locust Research Station / Jeddah”, “Abha / Nomas LT [light
trap?]”, 8.06.1972, }; Mecca, 10.01.1935, } (Philby) (BMNH); Senegal: Thie´s, 5.07.1908, } (“Eupyrgota
caffra Hend. 1913 }”) (Riggenbach) (ZMHB); Bandia, 18.06.1980, }, 9.07.1980, } (Sigwalt); M’Bour,
9.09.1980, {, } (Sigwalt) (MNHNP); Sudan: Ed Damer, Hudelba, 14.01, 5.03, 12.05.1962, 3 } (Remane)
(ZSSM); Khartoum, found on window, "Ent. coll. C11391", 2.11.1931, { (A. H. Wood); Shendi, at light,
"Africa 1/250,000 450 Map", "Ent. coll. 06856", 18.09.1928, } (Cowland) (NHMW); Togo: Lome, 9.11.1909
(NHMW); Uganda: “Ruwenzori Range B. M. E. Afr. Exp.”, “Kilembe / 4.500 ft”, 12.1934–01.1935, {, }
(Edwards) (BMNH); “Ankole / Kichwamba”, 1–5.05.1968, } (“Adapsilia latipennis (Walk.) d. G. Steyskal
72”) (Spangler) (USNM); “Kampala”, 19.07.1948, } (Sevastopulo) (NMKE); “Tororo / nesting pond [?]
/ early Am [?] [pencil, handwritten]”, 22.01.1959, } (collector unknown); Labwor 04.1955, } (van Someren)
(BMNH); Acholi: Madi Opei, 04.1951, } (Jackson) (NMKE); Yemen: Ta’izz, light trap, 26–28.05.1998, {,
19A Revision of Afrotropical Species of the Eupyrgota (Diptera, Pyrgotidae)...
3 } (van Harten et Awad); light trap, 26–28.05.1998, {, } (van Harten et Awad) (MHNG); light trap,
3–24.01.1999, } (van Harten et M. Mahyoub); 07.1999, {, }; 08.1999, }; 11.1999, }; 12.1999 (van Harten et
Awad) (ZMAN), 12.1999, } (van Harten et Awad) (MHNG); 05–06.2000, 8 }; 09.2001, 3 }; 09.2001, } (van
Harten et Al Yarimi) (ZMAN); Zimbabwe: “A. Carnegie / Dec. 1950 / Banket. S. R.”, } (Carnegie)
(NMPM); “Salisbury Exp. Stn. / Light Trap”, 11.1957, { (Brown) (SANC).
Non-type materia l possibly belonging to E. latipennis. Côte-d’Ivoire: “Lamto (Toumodi)”, 06–
08.1968, } (Girard) (MNHNP) (oviscape lacking; parafacials almost smooth, subshining; body and legs two-
colored); Uganda: Labwor Hills”, “Com. Inst. Ent. / Coll. No. 13261”, 04.1951, } (DEI; seen in 2002; not
re-examined); “Aequatorial-Afrika, Ruwenzori, Mubecki-River”, } (Hendel said it to be in BMNH; not loca-
ted; not examined); Zimbabwe: “Salisbury” [Harare], 8.01.1942, { (Pinhey) (NMPM).
20 V. A. Korneyev
Fig. 7. Eupyrgota latipennis: holotype Oxycephala latipennis } (BMNH) (1–3, 7, 9, 11), holotype } Adapsilia
ypsilon (BMNH) (4–6, 12) and non-type } from Yemen (ZMAN) (8, 10): 1, 4 — habitus (1 — dorsal, 4 —
left); 2, 5 — labels; 3, 6 — mesonotum, dorsal; 7, 9 — head (7 — left, 9 — anterior); 8 — parafacial; 10 —
foretrochanter; 11 — midfemur anteroventral; 12 — medial portion of R4+5 with setulae on dorsal side (1–7,
9, 11–12 — from dissecting microscope; 8, 10 — SEM).
Ðèñ. 7. Eupyrgota latipennis: ãîëîòèï Oxycephala latipennis } (BMNH) (1–3, 7, 9, 11), ãîëîòèï } Adapsilia
ypsilon (BMNH) (4–6, 12) è íåòèïîâàÿ } èç Éåìåíà (ZMAN) (8, 10): 1, 4 — îáùèé âèä (1 —
äîðñàëüíî, 4 — ñëåâà); 2, 5 — ýòèêåòêè; 3, 6 — ñðåäíåñïèíêà, äîðñàëüíî; 7, 9 — ãîëîâà (7 — ñëåâà,
9 — ñïåðåäè); 8 — ñêóëà; 10 — ïåðåäíèé âåðòëóã; 11 — ñðåäíåå áåäðî ïåðåäíåâåíòðàëüíî; 12 —
ñðåäíèé ó÷àñòîê R4+5 ñî ùåòèíêàìè íà äîðñàëüíîé ñòîðîíå (1–7, 9, 11–12 — ôîòî ñ áèíîêóëÿðà; 8,
10 — ôîòî ñî ñêàíèðóþùåãî ýëåêòðîííîãî ìèêðîñêîïà).
Dia gno s i s. Frontal plate and parafacial pitted; scutellum with 2–3 pairs of setae;
foretrochanter with at most 10–15 thickened setulae, forefemur slightly thickened, mid-
femur without femoral organ; oviscape on ventral side anterior to ventral sensillar lobe
and hooks with 4–5 brown thickened setulae on each side (fig. 8, 1: “s”; fig. 8, 2–3).
Similar to E. caffra Hendel in the frontal plate and parafacial pitted in both sexes and
foretrochanter sparsely spinulose in female, differing by the absence of femoral organ
and asymmetrical sclerites of praeputium in male. Very similar to E. sublatipennis in
having forefemur narrow, non-thickened in both sexes, asymmetrical sclerites of
praeputium in male and absence of the femoral organ in female, differing by the frontal
plate and dorsal half of the parafacial smooth, without expressed pits or wrinkles in both
sexes, and sparsely spinulose foretrochanter and shorter and wider aculeus in female.
21A Revision of Afrotropical Species of the Eupyrgota (Diptera, Pyrgotidae)...
Fig. 8. Eupyrgota latipennis, non-type } (1–5) and { (6–10) from Yemen (ZMAN): 1 — oviscape; 2 — apex
of oviscape, enlarged; 3 — group of thickened setulae anterior to hook, enlarged; 4 — spermathecae; 5 —
aculeus, ventral; 6 — male terminalia, right (segments 6–8 and phallus detached); 7 — medial and lateral
surstyli, posterior; 8 — proctiger, ventral; 9–10 — glans (9 — right, 10 — ventral).
Ðèñ. 8. Eupyrgota latipennis, íåòèïîâàÿ } (1–5) è { èç Éåìåíà (6–10) (ZMAN): 1 — îñíîâíîé ÷ëåíèê
ÿéöåêëàäà; 2 — âåðøèíà îñíîâíîãî ÷ëåíèêà ÿéöåêëàäà, óâåëè÷åíî, 3 — ãðóïïà óòîëùåííûõ ùåòèíîê
ñïåðåäè îò êðþêà, óâåëè÷åíî; 4 — ñïåðìàòåêè; 5 — ëåçâèå ÿéöåêëàäà, âåíòðàëüíî; 6 — òåðìèíàëèè
ñàìöà, ñïðàâà (ñåãìåíòû 6–8 è ôàëëþñ óäàëåíû); 7 — ñóðñòèëè, ñçàäè; 8 — ïðîêòèãåð, âåíòðàëüíî;
9–10 — ãëàíñ (9 — ñïðàâà, 10 — âåíòðàëüíî).
E. latipennis differs from both E. caffra and E. sublatipennis by having dark thickened
setulae on oviscape anterior to hooks in female.
R ed e s c r i p t i o n. Body yellow to reddish brown, usually without contrasting
brown patterns, in darker specimens with yellow scutellum and crossband in middle of
syntergite 1+2; setae brown to black, setulae reddish brown; mesonotum length
2.5–4.5 mm, wing length 9.1–13.0 mm in male, 9.3–13.5 mm in female (12.0 mm in
holotype).
Head (fig. 7, 7–9) pale yellow to light brown, eye ratio 0.52–0.56, genal-eye ratio
0.22–0.30; first flagellomere-pedicel ratio 1.18–1.33; medial vertical seta 0.28 times as
long as longest diameter of eye; lateral vertical seta 0.4–0.7 times as long as medial ver-
tical seta, often indistinguishable from surrounding setulae; postocellar seta (sometimes
2 pairs) 0.7–0.9 times as long as medial vertical seta; ocellar seta (sometimes 2 pairs)
as long as medial vertical seta; orbital seta 0.4–0.55 times as long as medial vertical
seta; frons yellow; frontal vitta matt yellow with sparse setulae; frontal plate and parafa-
cial matt or slightly subshining, shallowly pitted, with tiny yellowish brown setula in
bottom of every pit; rarely dorsal part of parafacial and frontal plate wrinkled; antenna
yellow; pedicel rather densely reddish brown setulose; first flagellomere microtrichose,
2.75 times as long as wide; face and frontogenal suture like in E. caffra; median carina
0.7–0.8 times as long as facial height; parafacial 1.4–2.0 times as wide as 1st flagellom-
ere; gena with or without dark subocular spot; postgena brownish yellow; occiput yel-
low; median occipital sclerite with 2–4 pairs of postocellar setulae in holotype and
9–10 setulae at each side, often with two oblique brown marks in the middle, forming
V- or Y-shaped mark (e. g., in holotype A. ypsilon); mouthparts as in E. caffra.
Thorax pale reddish yellow to brownish yellow, with notopleural triangle, anater-
gite and scutellum usually yellow; 2–5 pairs of scutellar setae (in holotype O. latipen-
nis, 3 on left and 2 on right side, in holotype A. ypsilon, 2 pairs).
Legs uniformly reddish yellow; forecoxa with 12–15 fine setulae and 1–2 stronger
setae; midcoxa with 1 strong seta on dorso-lateral part and row of 6–7 strong unmo-
dified setulae 0.9–1.0 times as long as longest seta on anteroventral margin; hindcoxa
with 1 strong seta on dorsolateral part, and 20–25 strong, posteroventrally directed setae
on anteroventral margin 0.7–0.9 times as long as seta, forming no comb; foretrochanter
with 6–7 sparse, but short spine-like setulae (fig. 7, 10); midtrochanter with 17–18 si-
milar setulae; hindtrochanter fine and sparsely setulose; all femora with subequal
1 basiventral seta, 1 slightly shorter seta at basiventral one-third of femora and with
thickened short setae ventrally, forming subequal anteroventral and posteroventral rows
in apical half; forefemur neither wrinkled, nor thickened, 4.2–4.7 times as long as wide
in female, and 4.5–4.8 in male.
Wing as in E. caffra; R4+5 bare in holotype O. latipennis and with 4–5 setulae
basally to r-m crossvein in holotype A. ypsilon; wing-thorax ratio 3.2–3.6, vein R4+5
ratio 1.4–1.55, vein M ratio 0.5.
Female abdomen reddish yellow to reddish brown, similar to E. caffra, but usual-
ly paler; syntergite 1+2 1.2–1.4 times as long as tergites 3–6 combined; oviscape sub-
shining reddish to brownish yellow, robust, 0.5–0.6 times as long as preabdominal ter-
gites combined and slightly longer than wide on ventral side; hooks simple, like in
E. caffra; ventral surface of oviscape anterior to hooks with 6–8 thickened dark brown
setulae (fig. 8, 2–3); aculeus with base gradually transiting into long stiletto-like pierc-
ing part more than 2.0–2.5 times as long as its bulky base (fig. 8, 5); spermathecae oval
(fig. 8, 4), 1.8–2.6 times as long as wide.
Male abdomen similarly colored; syntergite 1+2 1.1–1.3 times as long as tergites
3 and 4 combined; and 1.7–2.0 times as long as tergite 5; synsternite 1+2 5.2–6.0 times
as long as wide at posterior margin; epandrium and hypandrium as in fig. 8, 6; glans of
phallus with strongly asymmetrical sclerites of praeputium (fig. 8, 9–10), similar to
those in E. wagae Bigot (Korneyev, 2004: fig. 16, 4).
22 V. A. Korneyev
Rema r k s. E. latipennis is one of the most common and widespread pyrgotid
species in the Old World. It reaches southern Africa, where all the three species of the
group coexist. As these species were originally described and recognized from females,
the problem of identity of males was solved by study of males of E. latipennis from
Yemen, where there are no other Eupyrgota species. In this way, males of E. latipennis
and E. caffra were separated; however, noticeable differences in morphology of male
genitalia of E. latipennis and E. sublatipennis have not been found yet.
E. latipennis together with E. sublatipennis, and E. wagae from western Palaearctic
Asia belong to the same lineage in the latipennis group; its monophyly is supported by
having asymmetric praeputium sclerites, which are not known elsewhere in the family.
This group may include some other Asian species in which the morphology of the phal-
lus has not been studied properly.
Eupyrgota sublatipennis (Brunetti, 1929) (fig. 9)
Adapsilia sublatipennis Brunetti, 1929: 24; Adapsilia (Eupyrgota) sublatipennis: Steyskal, 1980: 557.
T yp e ma t e r i a l. Holotype } Adapsilia sublatipennis: Zimbabwe: “Hope Fountain / S. Rhodesia / (leg.
N. Jones) / 2.12.1921 / Rhodesia Museum”, “Type [red bordered circle]”, “Holo / type [red bordered
circle]”, “Adapsilia / sublatipennis / Brun Type } / Det. E. Brunetti 1926”, “92 [red ink]”, “Holotype
Adapsilia latipennis Brunetti verified J. E. Chainey 2002”, “BMNH # 252168” (BMNH).
Non - t y p e ma t e r i a l. South Africa: “Minastone [h/w illegible] / 16–15[sic!].2.1919 / H. G. Breyer”,
} (Breyer) (NMPM); Northern Province: “Zoutpan, Zpbg. [Zoutpansberg]”, 15–30.11.1932, } (van Son)
(NMPM); North West Province: “Mogol Nature Reserve / Ellisras Distr.”, 23°58' S / 27°45' E,
19–23.11.1979, 3 } (van Tonder); same locality, 19–23.11.1979, 3 } (C. Kok) (SANC); Mpumalanga:
Montrose Falls, 25°25' S 30°44' E, collected at light, 29.01.1989, } (Oberprieler) (SANC); Gauteng: Pretoria,
28.12.1915, } (Munro) (NMPM); Zimbabwe: “Hillside [Harare?], S. Rhod.”, 11.11.1922, } (no oviscape
ventroapical thickened setae; no femoral organ; pale wing; thin femora; foretrochanter without brush!
(uncommon for sublatipennis) (Swinbourne et Stevenson) (NMPM); Bulawayo, } (L. G. [?] Arnold [?]);
same locality, 8.12.1924, { (Stewensol [sic!]) (NMPM).
D i a gno s i s. Frontal plate neither pitted, nor wrinkled, matt; parafacial matt,
smooth, slightly wrinkled in ventral one-quarter; scutellum with 2–3 pairs of setae; of
40–60 thickened, hook-like setulae, forefemur not thickened, midfemur without
femoral organ; oviscape on ventral side anterior to ventral sensillar lobe and hooks with
4–5 thin yellowish setulae on each side. Differs from both E. latipennis and E. caffra
by having foretrochanter with dense brush in female and almost smooth frontal plate
and parafacial. Similar to E. latipennis in having forefemur narrow, non-thickened in
both sexes, asymmetrical sclerites of praeputium in male and absence of femoral organ
in female, differing by frontal plate and dorsal half of parafacial smooth, in both sexes,
thin yellowish setulae anterior to oviscape sensillar lobes, and shorter and wider aculeus
in female. E. sublatipennis differs from E. caffra by having asymmetrical sclerites of
praeputium in male and absence of femoral organ in female.
R ed e s c r i p t i o n. Body yellow to reddish brown, without contrasting brown pat-
terns, in darker specimens with scutellum, middle of syntergite 1+2 and posterior mar-
gins of tergites 4–6 yellow; setae brown to black, setulae reddish brown; mesonotum
length 2.3–3.5 mm (3.0 mm in holotype), wing length 7.5 mm in male, 8.2–10.2 mm
in female (8.9 mm in holotype).
Head (fig. 9, 3, 4) pale yellow, eye ratio 0.50–0.56, genal-eye ratio 0.21–0.33; first
flagellomere-pedicel ratio 1.0–1.27; frons yellow; frontal vitta, frontal plate and parafa-
cial matt smooth or with very small and shallow pits (fig. 9, 4, 5); antenna yellow; first
flagellomere 2.5–2.7 times as long as wide; face like in E. caffra; median carina
0.75–0.82 times as long as facial height; parafacial 1.8–2.0 times as wide as 1st flagel-
lomere; occiput yellow, sometimes with pale brown marks.
Thorax coloration as in E. latipennis; 2–3 pairs of scutellar setae.
Legs as in E. latipennis; foretrochanter mostly with 40–60 thickened, hook-like
setulae forming brush (fig. 9, 7, 8) in female, sparsely setulose in male and in aberrant
23A Revision of Afrotropical Species of the Eupyrgota (Diptera, Pyrgotidae)...
24 V. A. Korneyev
Fig. 9. Eupyrgota sublatipennis, holotype } (BMNH) (1–3, 6) and non-type } from Zimbabwe (NMPM) (4,
5, 7–15): 1 — habitus, left; 2 — labels; 3, 4 — head (3 — left, 4 — left anterior); 5 — parafacial; 6 — wing;
7 — foretrochanter, enlarged; 8 — foretrochanter and forefemur posteroventral; 9 — apex of oviscape; 10 —
group of thickened setulae anterior to hook, enlarged; 11 — spermathecae; 12 — aculeus, ventral; 13 — male
terminalia, right (segments 6–8, hypandrium and phallus detached); 14–15 — glans (14 — right, 15 —
ventral) (1–3, 6 — from dissecting microscope; 4–5, 7–10 — SEM; 11–15 — from compound microscope).
Ðèñ. 9. Eupyrgota sublatipennis, ãîëîòèï } (BMNH) (1–3, 6) è íåòèïîâàÿ } èç Çèìáàáâå (NMPM) (4,
5, 7–15): 1 — îáùèé âèä, ñëåâà; 2 — ýòèêåòêè; 3, 4 — ãîëîâà (3 — ñëåâà, 4 — ñëåâà ñïåðåäè); 5 —
ñêóëà; 6 — êðûëî; 7 — ïåðåäíèé âåðòëóã, óâåëè÷åíî; 8 — ïåðåäíèé âåðòëóã è ïåðåäíåå áåäðî çàäíå-
âåíòðàëüíî; 9 — âåðøèíà îñíîâíîãî ÷ëåíèêà ÿéöåêëàäà; 10 — ãðóïïà óòîëùåííûõ ùåòèíîê ñïåðåäè
îò êðþêà, óâåëè÷åíî; 11 — ñïåðìàòåêè; 12 — ëåçâèå ÿéöåêëàäà, âåíòðàëüíî; 13 — òåðìèíàëèè ñàìöà,
ñïðàâà (6–8-é ñåãìåíòû, ãèïàíäðèé è ôàëëþñ îòäåëåíû); 14–15 — ãëàíñ (14 — ñïðàâà, 15 —
âåíòðàëüíî) (1–3, 6 — ôîòî ñ áèíîêóëÿðà; 4–5, 7–10 — ôîòî ñî ñêàíèðóþùåãî ýëåêòðîííîãî
ìèêðîñêîïà; 11–15 — ôîòî ñî ñâåòîâîãî ìèêðîñêîïà).
female from Hillside (Zimbabwe); midtrochanter with 20–25 similar, but thinner and
sparser setulae in female; hindtrochanter and femora as in E. latipennis.
Wing as in E. latipennis, subapical spot usually pale brown, pterostigma and area;
R4+5 bare; wing-thorax ratio 3.0–3.5, vein R4+5 ratio 1.5–1.7, vein M ratio 0.6.
Female abdomen reddish to brownish yellow, similar to E. latipennis; syntergite
1+2 as in E. latipennis; oviscape subshining reddish yellow, 0.58–0.60 times as long as
preabdominal tergites combined and 1.10–1.15 as long as wide on ventral side; hooks
simple, or bilobate; ventral surface of oviscape anterior to hooks with 6–12 thin yellow-
ish setulae; aculeus with short piercing part 1.4–1.8 times as long as its bulky base; sper-
mathecae oval (fig. 9, 11), 1.3–1.6 times as long as wide.
Male abdomen brownish yellow; syntergite 1+2 with yellow crossband at middle,
1.3 times as long as tergites 3 and 4 combined; and 2.2 times as long as tergite 5; syn-
sternite 1+2 4.8 times as long as wide at posterior margin; epandrium and hypandrium
not dissected; glans of phallus with asymmetrical sclerites of praeputium.
Rema r k s. E. sublatipennis is restricted in its distribution to southeastern Africa.
It occurs sympatrically with E. caffra, but not with the widespread E. latipennis, which
only reaches Namibia. From the available material, it is not clear if E. sublatipennis
actually represents only a subspecies of E. latipennis.
Acze´l M. Revisio´n parcial de las Pyrgotidae neotropicales y anta ´rcticas, con sinopsis de los ge ´neros y especies
(Diptera, Acalyptratae) // Rev. Brasil. Ent. — 1956 a. — N 4. — P. 161–184.
Acze´l M. Revisio´n parcial de las Pyrgotidae neotropicales y anta ´rcticas, con sinopsis de los ge ´neros y especies
(Diptera, Acalyptratae) // Rev. Brasil. Ent. — 1956 b. — N 5. — P. 1–70.
Acze´l M. Revisio´n parcial de las Pyrgotidae neotropicales y anta ´rcticas, con sinopsis de los ge ´neros y especies
(Diptera, Acalyptratae) // Rev. Brasil. Ent. — 1956 c. — N 6. — P. 1–38.
Brunetti E. I. New African Diptera // Ann. Mag. Nat. Hist. — 1929. — (10) 4. — P. 1–35.
Coquillett C. D. Report on a collection of Japanese Diptera, presented to the U.S. National Museum by the
Imperial University of Tokyo // Proceedings of the United States National Museum. — 1898. — 21 [=
N 1146]. — P. 301–340.
Enderlein G. Klassifikation der Pyrgotiden // Sitzungsberichte der Gesellschaft Naturforschenden Freunde zu
Berlin. — 1942 (1941). — H. 2. — S. 98–134.
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H. 11. — S. 77–117 + 1 Taf.
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// Vestn. zoologii. — 2004. — 38, N 1. — P. 19–46.
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Stuttgart. Beitr. Naturk. — 1972. — N 238. — P. 1–11.
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25A Revision of Afrotropical Species of the Eupyrgota (Diptera, Pyrgotidae)...
|
| id | nasplib_isofts_kiev_ua-123456789-9418 |
| institution | Digital Library of Periodicals of National Academy of Sciences of Ukraine |
| issn | 0084-5604 |
| language | English |
| last_indexed | 2025-12-07T17:51:26Z |
| publishDate | 2006 |
| publisher | Інститут зоології ім. І.І. Шмальгаузена НАН України |
| record_format | dspace |
| spelling | Korneyev, V.A. 2010-06-30T16:14:28Z 2010-06-30T16:14:28Z 2006 A revision of afrotropical species of the eupyrgota (diptera, pyrgotidae): the spinifemur group and latipennis subgroup of species / V.A. Korneyev // Вестн. зоологии. — 2006. — Т. 40, № 1. — С. 3-25. — Бібліогр.: 18 назв. — англ. 0084-5604 https://nasplib.isofts.kiev.ua/handle/123456789/9418 595.773.4 Based on the study of the type specimens described
 by F. Walker, F. Hendel, E. Brunetti and P. Vanschuytbroeck and exhaustive material from many
 entomological collections, species of the spinifemur group and latipennis subgroup of the genus
 Eupyrgota Coquillett occurring in the Afrotropical Region and Arabian Peninsula are redescribed. The
 following synonymy is established. Eupyrgota latipennis (Walker, 1849), comb. n. (= Oxycephala
 latipennis Walker) = Adapsilia ypsilon Hendel, 1914, syn. n. Eupyrgota spinifemur (Hendel, 1934) comb. n.
 (= Adapsilia spinifemur Hendel) = Metropina nigra Vanschuytbroeck, 1963, syn. n. Eupyrgota saegeri
 (Vanschuytbroeck, 1963) comb. n. is transferred from Lygiohypotyphla Hendel. На основании изучения типовых материалов, описанных Ф. Уокером, Ф. Генделем, Ю. Брунетти и П. Вансхётбруком и исчерпывающих материалов из многочисленных энтомологических коллекций, представлены описания видов из группы spinifemur и подгруппы latipennis рода Eupyrgota, встречающихся в Афротропической области и на Аравийском полуострове. Установлена следующая синонимия. Eupyrgota latipennis (Walker, 1849), comb. n. (= Oxycephala latipennis Walker) = Adapsilia ypsilon Hendel, 1914, syn. n. Eupyrgota spinifemur (Hendel, 1934) comb. n. (= Adapsilia spinifemur Hendel) = Metropina nigra Vanschuy-tbroeck, 1963, syn. n. Eupyrgota saegeri (Vanschuytbroeck, 1963) comb. n. перемещена из Lygiohypo-typhla Hendel. en Інститут зоології ім. І.І. Шмальгаузена НАН України Фауна и систематика A revision of afrotropical species of the eupyrgota (diptera, pyrgotidae): the spinifemur group and latipennis subgroup of species Ревизия афротропических видов рода Eupyrgota (Diptera, Pyrgotidae): группа видов spinifemur и подгруппа latipennis Article published earlier |
| spellingShingle | A revision of afrotropical species of the eupyrgota (diptera, pyrgotidae): the spinifemur group and latipennis subgroup of species Korneyev, V.A. Фауна и систематика |
| title | A revision of afrotropical species of the eupyrgota (diptera, pyrgotidae): the spinifemur group and latipennis subgroup of species |
| title_alt | Ревизия афротропических видов рода Eupyrgota (Diptera, Pyrgotidae): группа видов spinifemur и подгруппа latipennis |
| title_full | A revision of afrotropical species of the eupyrgota (diptera, pyrgotidae): the spinifemur group and latipennis subgroup of species |
| title_fullStr | A revision of afrotropical species of the eupyrgota (diptera, pyrgotidae): the spinifemur group and latipennis subgroup of species |
| title_full_unstemmed | A revision of afrotropical species of the eupyrgota (diptera, pyrgotidae): the spinifemur group and latipennis subgroup of species |
| title_short | A revision of afrotropical species of the eupyrgota (diptera, pyrgotidae): the spinifemur group and latipennis subgroup of species |
| title_sort | revision of afrotropical species of the eupyrgota (diptera, pyrgotidae): the spinifemur group and latipennis subgroup of species |
| topic | Фауна и систематика |
| topic_facet | Фауна и систематика |
| url | https://nasplib.isofts.kiev.ua/handle/123456789/9418 |
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