A Review of the Subgenus Chilothorax of the Genus Aphodius (Coleoptera, Scarabaeidae) Occurring in Mongolia, with Description of New Species
The Mongolian species of Aphodius Illiger belonging to the subgenus Chilothorax Motschulsky are reviewed. A. kerzhneri Nikolajev and A. mongolaltaicus Nikolajev are placed in the subgenus Chilothorax. Two new synonyms (A. altaicus Nikolajev = A. puntsagdulamae Nikolajev...
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Інститут зоології ім. І.І. Шмальгаузена НАН України
2001
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| Cite this: | A Review of the Subgenus Chilothorax of the Genus Aphodius (Coleoptera, Scarabaeidae) Occurring in Mongolia, with Description of New Species / A.V. Frolov // Вестн. зоологии. — 2001. — Т. 35, № 3. — С. 39-45. — Бібліогр.: 1 назв. — англ. |
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| citation_txt | A Review of the Subgenus Chilothorax of the Genus Aphodius (Coleoptera, Scarabaeidae) Occurring in Mongolia, with Description of New Species / A.V. Frolov // Вестн. зоологии. — 2001. — Т. 35, № 3. — С. 39-45. — Бібліогр.: 1 назв. — англ. |
| collection | DSpace DC |
| description | The Mongolian species of Aphodius Illiger
belonging to the subgenus Chilothorax Motschulsky are reviewed. A. kerzhneri Nikolajev and
A. mongolaltaicus Nikolajev are placed in the subgenus Chilothorax. Two new synonyms (A. altaicus
Nikolajev = A. puntsagdulamae Nikolajev, syn. n.; A. grafi Reitter = A. boreomongolicus Nikolajev,
syn. n.) are established. Three new species are described. A key to Mongolian species of Chilothorax is
presented.
Дан обзор обитающих в Монголии видов подрода Chilothorax Motschulsky рода Aphodius Illiger. В этот подрод помещены Л. kerzhneri Nikolajev и Л. mongolaltaicus Nikolajev. Установлена синонимия двух видовых названий (Л. altaicus Nikolajev = A. puntsagdulamae Nikolajev, syn. n.; A. grafi Reitter = A. boreomongolicus Nikolajev, syn. n.). Описаны 3 новых вида. Приведена таблица для определения монгольских видов подрода Chilothorax.
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Vestnik zoologii, 35(3): 39—45, 2001
© 2001 A. V. Frolov
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17327
UDC 595.764.1(517.3)
A REVIEW OF THE SUBGENUS CHILOTHORAX OF THE GENUS
APHODIUS (COLEOPTERA, SCARABAEIDAE) OCCURRING
IN MONGOLIA, WITH DESCRIPTION OF NEW SPECIES
A. V. Frolov
Institute of Zoology NAS of Belarus, Academicheskaya str., 27, Minsk, 220072 Belarus
E-mail: avfrolov@mail.ru.
Accepted 10 May 2000
A Review of the Subgenus Chilothorax of the Genus Aphodius (Coleoptera, Scarabaeidae) Occurring in
Mongolia, with Description of New Species. Frolov A. V. – The Mongolian species of Aphodius Illiger
belonging to the subgenus Chilothorax Motschulsky are reviewed. A. kerzhneri Nikolajev and
A. mongolaltaicus Nikolajev are placed in the subgenus Chilothorax. Two new synonyms (A. altaicus
Nikolajev = A. puntsagdulamae Nikolajev, syn. n.; A. grafi Reitter = A. boreomongolicus Nikolajev,
syn. n.) are established. Three new species are described. A key to Mongolian species of Chilothorax is
presented.
Key wo rd s : Coleoptera, Scarabaeidae, Aphodius, Chilothorax, taxonomy, new species, new syno-
nyms, Mongolia.
Îáçîð ïðåäñòàâèòåëåé ïîäðîäà Chilothorax ðîäà Aphodius (Coleoptera, Scarabaeidae) Ìîíãîëèè ñ
îïèñàíèåì íîâûõ âèäîâ. Ôðîëîâ À. Â. – Äàí îáçîð îáèòàþùèõ â Ìîíãîëèè âèäîâ ïîäðîäà
Chilothorax Motschulsky ðîäà Aphodius Illiger. Â ýòîò ïîäðîä ïîìåùåíû A. kerzhneri Nikolajev è
A. mongolaltaicus Nikolajev. Óñòàíîâëåíà ñèíîíèìèÿ äâóõ âèäîâûõ íàçâàíèé (A. altaicus Nikola-
jev = A. puntsagdulamae Nikolajev, syn. n.; A. grafi Reitter = A. boreomongolicus Nikolajev, syn. n.).
Îïèñàíû 3 íîâûõ âèäà. Ïðèâåäåíà òàáëèöà äëÿ îïðåäåëåíèÿ ìîíãîëüñêèõ âèäîâ ïîäðîäà
Chilothorax.
Êëþ÷åâûå ñëîâà : Coleoptera, Scarabaeidae, Aphodius, Chilothorax, òàêñîíîìèÿ, íîâûå âèäû,
íîâûå ñèíîíèìû, Ìîíãîëèÿ.
Introduction
The Mongolian fauna of Aphodius Illiger is not very rich, but does include a few apparently endemic
species. Thanks to several entomological expeditions taken to Mongolia, a rather extensive amount of
Aphodiinae material has been collected and deposited in several museums. Much of this material remains to
be examined.
In the present article, the Mongolian species belonging to the subgenus Chilothorax Motschulsky are
reviewed. This subgenus is very difficult taxonomically primarily because of great variability in diagnostic
characters, particularly elytral pattern, used by previous authors. I have included in this review some species
described by G. Nikolajev (Nikolajev, Puntsagdulam, 1984) from Mongolia as representatives of the subge-
nus Pseudacrossus Reitter, which belong to the subgenus Chilothorax. Three species are described as new.
Exact label data are cited in quotes for the type specimens. Author's remarks are in square brackets.
The following abbreviation identify the collections housing the material examined: DKCP – David
Král's collection, Czech Republic, Praha; MLUH – Martin-Luther-Universitat, Germany, Halle; NMPC –
National Museum (Natural History), Czech Republic, Praha; ZISP – Zoological Institute, Academy of
Sciences, Russia, St.-Petersburg.
Key to the Chilothorax species from Mongolia
Òàáëèöà äëÿ îïðåäåëåíèÿ âèäîâ ïîäðîäà Chilothorax èç Ìîíãîëèè
1. Eye relatively large: its diameter in dorsal view approximately two times longer then minimum
distance between eye margin and gula (fig. 14); fore tibia of male relatively slender, its inner
margin slightly convex (fig. 11); disc of metasternum not pubescent; elytral pattern consisting of
a few small brown maculae; apices of parameres without feebly sclerotized extensions (fig. 23). ...
...............................................................................................................A. mongolaltaicus Nikolajev
A. V. Frolov 40
– Eye relatively small: its diameter in dorsal view approximately as long as minimum distance
between eye margin and gula (fig. 15). ......................................................................................... 2
2. Disc of head and pronotum dark brown to nearly black. ............................................................. 3
– Disc of head and pronotum brown to light brown. ...................................................................... 5
3. Elytra entirely brown to dark brown; fore tibia relatively slender, its inner margin straight
(fig. 12); parameres (fig. 22). ...................................................................... A. kerzhneri Nikolajev
– Elytra light brown with a few small dark brown maculae, or immaculate; fore tibia not slender,
its inner margin concave. .............................................................................................................. 4
4. Longer (5.2—6.0 mm); apical spur of fore tibia relatively long in male (fig. 10); disc of metaster-
num not pubescent; parameres relatively long (fig. 16). ................................. A. altaicus Nikolajev
– Shorter (3.3—4.6 mm); apical spur of fore tibia relatively short (fig. 13); disc of metasternum
pubescent in male; parameres relatively short (fig. 24). ...................................... A. comma Reitter
5. Metasternum not pubescent; parameres with long extensions (fig. 19). ...... A. jacobsoni W. Kosh.
– Metasternum pubescent. ............................................................................................................... 6
6. Longer (6.4—6.7 mm); parameres (fig. 21). ............................ A. chandmanicus Frolov, sp. n. ({)
– Shorter (4.4—6.1 mm). ................................................................................................................. 7
7. Scutellum with hairs posteriorly (hairs are best viewed with oblique lighting, but may be abraded
in some individuals); parameres with long extensions (fig. 20). ................. A. alexis Frolov, sp. n.
– Scutellum without hairs posteriorly (only base of scutellum pubescent); parameres with short
extensions (fig. 17, 18). ................................................................................................................. 8
8. Apices of parameres more slender in lateral view (fig. 18); genae in females not protruding be-
yond eyes. ............................................................................................... A. tanhensis Frolov, sp. n.
– Apices of parameres less slender in lateral view (fig. 17); genae in females protruding beyond
eyes. .......................................................................................................................... A. grafi Reitter
A. (Chilothorax) altaicus Nikolajev, 1984 (fig. 1, 10, 16)
= A. gobiensis Balthasar, 1967 (nom. praeocc.) — A. puntsagdulamae Nikolajev, 1984, syn. n.
M a t e r i a l e x a m i n e d . Holotype { labeled "Gobi-Altai, Ich Bogd, 2.VI.62, leg. Grebenscikov",
"Aphodius (Volinus) gobiensis n. sp. {, 66, Balthasar, Holotypus", "A. jacobsoni Kosh., det. dr. Endrodi
1968", "Aphodius jacobsoni gobiensis Blt., det. Grebenscikov 1971"; 6 paratypes { with the same data. One
paratype in NMPC, holotype and other paratypes in MLUH.
A. puntsagdulamae. Holotype { labeled "Á.-Óëã. ð. Èõ-Äæàðàëòûí-Ãîë, 20 êì ÑÇ Áóëãàíà [Bajan-
Ulgij aimak, Ikh-Dzharaltyn-Gol river, 20 km NW from Bulgan] 4.VII.1980, Ïóíöàãäóëàì [Puntsagdulam
leg.]" (ZISP); paratypes: } with the same data and { labeled «Êîáä. ð. Äýä-Íàðèéí-Ñàëà, ïðèòîê ð. Áóë-
ãàíà [Hovd aimak, Ded-Narijn-Sala river, tributary of Bulgan river], 3.VII.1980, Ïóíöàãäóëàì» (ZISP).
Gov'-Altaj: Tegreg, 19.06.1980, 3 spm. (Medvedev, ZISP).
D i f f e r e n t i a l d i a g n o s i s . The species is very similar to A. nigrivittis Solsky,
occurring in Central Asia and Kazakhstan; can be separated from it by paramere shape
and elytral pattern.
D i s t r i b u t i o n . Mongolia.
A. (Chilothorax) kerzhneri Nikolajev, 1984 (fig. 2, 12, 22)
M a t e r i a l e x a m i n e d . Holotype { labeled «Ìîíãîëèÿ, Àðà-Õàíãàéñêèé àéì. [Mongolia,
Arhangaj aimak], ð. Õîéò-Òåðèí-Ãîë, 35 êì ÞÇ Öàõèðà [Hoit-Terhin-Gol riv., 35 km SW from Tsahir],
27—28.VI.1975, Ñóãîíÿåâ [Sugonyaev leg.]» (ZISP); 5 paratypes: { and } with the same data, and 3 { la-
beled «Ìîíãîëèÿ, Àðà-Õàíãàéñêèé àéì., [Arhangaj aimak], 7 êì ÞÇ Òàðÿòà [7 km SW from Taryat],
24—26.VI.1975, Ãóðüåâà [Gur'eva leg.]» (ZISP).
D i f f e r e n t i a l d i a g n o s i s . The species is similar to A. praenubilis Balth. (dis-
tributed in Tian-Shan mts.) and particularly to A. exilimanus Kabakov, recently de-
scribed from European Russia (Ul'yanovsk Prov.); differing from both species by the
entirely dark elytra and, additionally, from the first by the denser punctation of the
pronotum and head.
D i s t r i b u t i o n . Mongolia: Arhangaj, Dzabhan, Bajan-Ulgij.
N o t e . The species was originally described as relating to subgenus Pseudacrossus.
Known only from the type material.
Description of New Species of the Genus Aphodius … 41
A. (Chilothorax) mongolaltaicus Nikolajev, 1984 (fig. 3, 11, 14, 23)
M a t e r i a l e x a m i n e d . Holotype { labeled «ÌÍÐ [Mongolia], Þ. áåð. îç. Äóíä-Íóð [south
shore of Dund Nuur lake], îêð. Öàãàí-Íóðà [near Cagaannuur], 2.07.1978, Ãóðüåâà [Gurjeva leg.]»
(ZISP); paratype } labeled «ÌÍÐ, ð. Äçàáõàí, 45 êì Ñ. Ç. Äýëãåðà [Dzabkhan riv., 45 km NW from Del-
ger], 23.8. 1978, Ãóðüåâà» (ZISP).
Bajan-Ulgij: Tolbo-nuur, 1.07.1968, 1 spm. (exp. Z. Kaszab, DKCP); Gov'-Altaj: 12 km E from
Jesonbulag, h=2220 m, 1 spm. (exp. Kaszab, DKCP).
D i f f e r e n t i a l d i a g n o s i s . The species differs from all Mongolian Chilotho-
rax species by relatively big eye, and by the relatively slender fore tibia with slightly
convex inner margin (except for A. kerzhneri Nikolajev).
D i s t r i b u t i o n . Mongolia, South-Western Siberia (Altai mts.).
N o t e . This species was originally described in the subgenus Pseudacrossus on the
basis of having the lower spur of middle tibia truncate in the male. However, examina-
tion of available male specimens, including the holotype, has shown that the lower
Fig. 1—9. Aphodius. Habitus of males: 1 – A. altaicus; 2 – A. kerzhneri; 3 – A. mongolaltaicus; 4 – A. alexis;
5 – A. chandmanicus; 6 – A. jacobsoni; 7 – A. grafi; 8 – A. comma; 9 – A. tanhensis.
Ðèñ. 1—9. Aphodius. Ãàáèòóñ ñàìöîâ: 1 – A. altaicus; 2 – A. kerzhneri; 3 – A. mongolaltaicus; 4 – A. alexis;
5 – A. chandmanicus; 6 – A. jacobsoni; 7 – A. grafi; 8 – A. comma; 9 – A. tanhensis.
A. V. Frolov 42
spurs of middle tibiae are acute, not truncate, and at least half as long as the upper
spur, or longer.
A. (Chilothorax) jacobsoni W. Koshantschikov, 1911 (fig. 6, 15, 19)
M a t e r i a l e x a m i n e d . Lectotype (here designated) { labeled «Ñ. Â. Ìîíãîëèÿ, õð. Õàíãàé
[North-Eastern Mongolia, Hangaj mts.], Ïîòàíèí [Potanin leg.]», "A. Jacobsoni v. Georgi m." (ZISP);
10 paralectotypes: 6 { and 4 } with the same data (ZISP).
Bajanhongor: Orog nuur [lake], 3—14.06.1926, 2 spm. (ZISP); Uvurhangai: Hoolt, 16—27.07.26,
1 spm. (ZISP); Umnegov: Dalanzadgad, 06.1964, 1 spm. (Muche, DKCP).
D i f f e r e n t i a l d i a g n o s i s . The species is similar to A. alexis Frolov, sp. n.
and A. grafi; differs from the first by the shape of the parameres, and by the glabrous
disc of metasternum; and from the second by the shape of the parameres and angulate
genae in female.
D i s t r i b u t i o n . Mongolia, Russia (Southern Siberia).
A. (Chilothorax) alexis Froliv, sp. n. (fig. 4, 20)
T y p e m a t e r i a l . Holotype { labeled «Ñ. Â. Ìîíãîëèÿ, õð. Õàíãàé [North-Eastern Mongolia,
Hangaj mts.], Ïîòàíèí [Potanin leg.]», "A. Jacobsoni, v. Alexis {, m."; 9 paratypes: 3 { with the same data;
4 { labeled «Ñ. Â. Ìîíãîëèÿ, õð. Õàíãàé, Ïîòàíèí», "A. Jacobsoni, m. {"; } labeled «Ñ. Â. Ìîíãîëèÿ,
õð. Õàíãàé, Ïîòàíèí», "A. Jacobsoni, v. Georgi }, m."; } labeled "Mongol. Changai geb. Potanin". All
specimens deposited in ZISP.
D e s c r i p t i o n . Holotype { (fig. 4). Head brown, shiny, punctures unequal,
relatively dense. Frons not tuberculate, frontal suture widely broken at the middle.
Clypeus very faintly sinuate at middle, widely rounded at sides. Genae angulate.
Pronotum brown, shiny, with unequal punctures, widely light brownish on sides; sides
and base bordered; hind angles rounded; lateral margins pubescent with sparse hairs.
Scutellum brown, shiny, punctured, with hairs posteriorly (hairs are best viewed with
oblique lighting). Elytra lightly brown, shiny, with brown spots on interstices 3—7.
Striae deep, with large punctures; interstices feebly convex, finely punctured. Sides and
apices of elytra finely covered with short hairs. Ventrum of body brown, femora and
tibiae slightly lighter. Disc of metasternum pubescent. Apical spur of fore tibia acute
and curved downward. Apical spurs of middle tibia slender and long; lower spur
slightly more then half the length of the upper. First segment of hind tarsi as long as
upper apical spur of tibia and a bit longer then two following segments together. Ad-
joining apical setae of hind tibiae short and equal except for some outermost ones. Pa-
rameres with auriculate extensions (fig. 20). Body length 5.4 mm.
Female can be separated from male by the narrower pronotum, denser punctures
on pronotum and head, and by the presence of finer pubescence on the elytral apices.
Body length of paratypes 4.6—5.5 mm.
D i f f e r e n t i a l d i a g n o s i s . The species is similar to A. jacobsoni and A. grafi;
differs from first by the paramere shape, and by the pubescent disc of metasternum;
and from second by angulate genae (female), and by the shape of parameres. From
both species, it may be separated by scutellum possessing short light recumbent hairs
posteriorly (the hairs may be abraded in some individuals and consequently this is
probably not a reliable diagnostic character).
E t y m o l o g y . The name of one variation, designated on the original labels by
W. Koshantschikov, was used as the name of the new species.
A. (Chilothorax) grafi Reitter, 1901 (fig. 7, 17)
= A. boreomongolicus Nikolajev, 1984, syn. n.
M a t e r i a l e x a m i n e d . A. boreomongolicus: holotype { and paratype { labeled "Uvs, Bukhmuren,
1978.VII.7, Puntsagdulam [leg.]" (ZISP).
Description of New Species of the Genus Aphodius … 43
Bajan-Hongor: Holbogiin-nur, 30 km N Orog-nuur [lake], 4.07.1973, 2 spm. (Medvedev); Uvs:
Bukhmuren, 1978, 1 spm.; Hovd: Zhargalant, 22.07.1926, 7 spm. (Kozlov); Umnegov: Huryn-Halha-nuru,
25 km W Nojon [Hovuun], 20.06.1973, 2 spm. (Medvedev); Hovsgol: Sumber, 13.07.1975 (Gur'eva). All
specimens deposited in ZISP.
D i f f e r e n t i a l d i a g n o s i s . The species is similar to A. jacobsoni W. Kosh.,
A. alexis sp. n., and A. tanhensis sp. n.; differs from the first by the pubescent disc of
metasternum in both sexes. Males of these four species can easily be separated by the
shape of parameres and their extensions. At best, females are difficult to identify.
D i s t r i b u t i o n . Russia (Altai mts.), Mongolia.
N o t e . The original description of A. boreomongolicus was based on the series of
three specimens from Bukhmuren, Uvs aimak. The species was conditionally placed in
the subgenus Nobius Muls. et Rey. Having examined these type specimens I could not
detect any essential characters, including in the shape of the parameres, which would
Fig. 10—24. Aphodius. 10—13 – fore tibiae of males: 10 – A. altaicus; 11 – A. mongolaltaicus; 12 –
A. kerzhneri; 13 – A. comma. 14, 15 – head in ventral view: 14 – A. mongolaltaicus; 15 – A. jacobsoni. 16—
24 – parameres in dorsal and lateral view: 16 – A. altaicus; 17 – A. grafi; 18 – A. tanhensis; 19 –
A. jacobsoni; 20 – A. alexis; 21 – A. chandmanicus; 22 – A. kerzhneri; 23 – A. mongolaltaicus; 24 –
A. comma.
Ðèñ. 10—24. Aphodius. 10—13 – ïåðåäíÿÿ ãîëåíü ñàìöîâ: 10 – A. altaicus; 11 – A. mongolaltaicus; 12 –
A. kerzhneri; 13 – A. comma. 14, 15 – ãîëîâà ñíèçó: 14 – A. mongolaltaicus, 15 – A. jacobsoni. 16—24 –
ïàðàìåðû ñâåðõó è ñáîêó: 16 – A. altaicus; 17 – A. grafi; 18 – A. tanhensis; 19 – A. jacobsoni; 20 – A. alexis;
21 – A. chandmanicus; 22 – A. kerzhneri; 23 – A. mongolaltaicus; 24 – A. comma.
A. V. Frolov 44
allow me to distinguish A. boreomongolicus from A. grafi, inhabiting the same regions of
Mongolia and contiguous territories of Russia. According to the original descrip-
tion (Nikolajev, Puntsagdulam, 1984: 155) the main character to distinguish
A. boreomon-golicus from similar species (A. grafi, A. kerzhneri, A. altaicus) was the
monotonous brown color of elytra. But having examined the large series of A. grafi
I found that the elytral pattern of this species vary greatly. In this context, I suppose
that the type specimens of A. boreomongolicus are in fact uncommonly melanized indi-
viduals of A. grafi.
A. (Chilothorax) tanhensis Frolov, sp. n. (fig. 9, 18)
T y p e m a t e r i a l . Holotype { labeled «Ìîíã. [Mongolia] 21.V.[18]94 Íàðàñíåé [Narasnej-Zakh,
Selenge aimak] Êø [Kashkarov leg.]», "Volinus nigrivittis Solsky, W. Koshantschikov det" (ZISP); 23 para-
types: { with the same data (ZISP); 5 { with the same data but without the labels of W. Koshantschi- kov
(ZISP); { and 14 } labeled «êë. Òàíõå, ñ.-ç. Ìîíã. [NW Mongolia, Uvs aimak, Tanhe spring, near Ulaan-
gom] 23.V.[19]23, Ñåâêî, Ñèçîâà [Sevko and Sizova leg.]» (ZISP); { and } labeled "Mongolia 27.6.[19]90
Ulaanbaatar airport D. Král lgt", "coll. D. Král" (DKCP).
D e s c r i p t i o n . Holotype { (fig. 9). Head dark brown, shiny, densely punctate.
Frons not tuberculate. Clypeus very feebly sinuate at middle, rounded at sides. Genae
small, angulate. Pronotum dark brown on disc, light brownish on sides, shiny, with
equal punctures; sides and base bordered; hind angles rounded; lateral margins pubes-
cent with sparse short hairs. Scutellum brown, densely punctured. Elytra yellowish
brown, shiny, with a pattern of longitudinal brown spots (of the same color or slightly
lighter then pronotal disc) on interstices 3—8. Striae fine; punctures of striae bigger in
diameter then width of the stria; interstices feebly convex, punctured. Sides and apices
of elytra finely pubescent with very short, almost invisible hairs. Ventrum of body
brown, femora and tibiae slightly lighter. Disc of metasternum slightly concave, pubes-
cent with dense hairs. Apical spur of fore tibia acute and curved downward. Apical
spurs of middle tibia slender and long; lower spur more then half the length of the up-
per. First segment of hind tarsi as long as upper spur of tibia and a bit longer then two
following segments together. Most of adjoining apical setae of hind tibiae short and of
equal length, but some setae relatively long. Parameres with short extensions, their api-
ces slender and acute (fig. 18). Body length 5.5 mm.
Female can be separated from male by the narrower pronotum, and denser punc-
tures on pronotum and head.
Body length of paratypes 5.2—5.6 mm.
D i f f e r e n t i a l d i a g n o s i s . The species is similar to A. grafi Reitt. The only
reliable character separating the two species is the shape of the parameres. In the series
examined, females of A. grafi had the genae more protruding then the eyes, whereas
females of A. tanhensis Frolov, sp. n. had the genae less protruding then the eyes; how-
ever, this character may prove to be unreliable.
D i s t r i b u t i o n . Mongolia. According to the figures of parameres of A. grafi in
the work of Nikolajev and Puntsagdulam (1984: 139), which quite correspond to the
parameres of A. tanhensis Frolov, sp. n., the data on distribution of A. grafi perhaps
belong, at least partly, to A. tanhensis Frolov, sp. n., and the latter is distributed also in
Hovd, Gov'-Altaj, and Bajanhongor aimaks.
E t y m o l o g y . The name of the new species was derived from the toponym
"Tanhe".
A. (Chilothorax) comma Reitter, 1892 (fig. 8, 13, 24)
M a t e r i a l e x a m i n e d . Tov: Songino, SW Ulaanbaator, 18.06.1967, 14 spm. (Zaitsev); northern
slope of Bogdo Uula mts., near Ulaanbaator, 15.06.1967, 1 spm., 29.06.1967, 1 spm. (Zaitsev), 29.06.1967,
1 spm. (Kerzhner); Ulaanbaator, 11.06.1967, 1 spm (Emel'yanov), 19.06.1967, 1 spm. (Kerzhner); Nalajh,
25.06.1967, 3 spm. (Kerzhner); Kerulen, 1.06.1976, 5 spm. (Kerzhner), 1 spm. (Gur'eva); Hovsgol: Achi-
mag nur, 25 km E Muren, 27.06.1968, 1 spm. (Kozlov); Uvs: Hyargas nuur, 3.07.1978, 1 spm. (Puntsagdu-
Description of New Species of the Genus Aphodius … 45
lam); Dornod: 15 km N Hoh nor, 28.06.1976, 4 spm. (Gur'eva); Herlen gol, near Hulen-Bujra, 3.06.1976,
6 spm. (Gur'eva); Galin gol, 33 km SE Halhgol, 10.06.1976, 6 spm. (Gur'eva); Numregin gol, 32 km SE
Salhid Ondor mts., 16.06.1976, 5 spm. (Gur'eva); Bulgan: 5 km W Bajannuur, 16.06.1976, 5 spm.
(Gur'eva); Tuul gol, Zaamryi uuliaas, 23.06.1978, 1 spm. (Puntsagdulam); Gov'-Altaj: 45 km SW Bajan,
17.06.1980, 1 spm. (Puntsagdulam); Dzabhan: Songino, 12.06.1980, 1 spm. (Puntsagdulam); Uverhangaj:
50 km W Arvajheer, 13.06.1980, 3 spm. (Medvedev); Ongijn Gol, 14.07.1926, 1 spm. (Kirichenko). All
specimens deposited in ZISP.
D i f f e r e n t i a l d i a g n o s i s . The species differs from all other Mongolian
Chilothorax species by the shape of parameres and the relatively small size.
D i s t r i b u t i o n . Northern Kazakhstan, Transbaical region, Mongolia.
A. (Chilothorax) chandmanicus Frolov, sp. n. (fig. 5, 21)
T y p e m a t e r i a l . Holotype { labeled «Ìîíãîëèÿ ÑÂ, ×àíäìàíü [North-Eastern Mongolia, Hovd
aimak, Chandman'], 10.VII.1980, Ã. Ìåäâåäåâ [G. Medvedev leg.]» (ZISP); 3 paratypes: 2 { with the same
data (ZISP); { labeled "Mongolia, Chovd aimak, Jamatin Dolon, cca 40 km N von Somon Manchan am
SW Ecke des Char us nuur 1200 m, Exp. Dr. Z. Kaszab, 1966", "Nr. 673, 11.VII.1966" (DKCP).
D e s c r i p t i o n . Holotype { (fig. 5). Head brown, shiny, sparsely punctured.
Frons not tuberculate. Clypeus sinuate at middle, rounded at sides. Genae angulate.
Pronotum brown, shiny, sparsely irregularly punctured, lightly brownish on sides; sides
and base bordered; hind angles rounded; lateral margins pubescent with long hairs.
Scutellum triangular, brown, shiny, with sparse punctures. Elytra lightly yellowish
brown, shiny, with longitudinal brown spots on interstices 3—7. Striae deep, with big
punctures; interstices feebly convex, punctured. Sides and apices of elytra finely pubes-
cent with minute hairs. Ventrum of body light brown. Disc of metasternum pubescent.
Fore tibia slender, its apical spur acute and curved downward. Apical spurs of middle
tibia slender and long; lower spur more then half the length of the upper. First seg-
ment of hind tarsi as long as or a bit longer then upper tibial spur and as long as two
following segments together. Adjoining apical setae of hind tibiae unequal, relatively
short. Parameres with extensions (fig. 21). Body length 6.7 mm.
Female unknown.
Body length of paratypes 6.4—6.7 mm.
D i f f e r e n t i a l d i a g n o s i s . The species differs from all Mongolian Chilotho-
rax species by its larger size and by paramere shape.
E t y m o l o g y . The name of the new species was derived from the toponym
"Chandman'".
Acknowledgements
I would like to thank Dr. Hans Fery (Berlin), Dr. Oleg Kabakov (St. Petersburg), Dr. Izyaslav
Kerzhner (St. Petersburg), Dr. David Král (Praha), Dr. Georgy Nikolajev (Almaty), Dr. Karla Schnei-
der (Halle), and Dr. Alexey Solodovnikov (St. Petersburg) for valuable help. Special thanks are due to
Dr. Paul Lago (University of Mississippi) for linguistic review of the manuscript and to Mr. Kirill Dow-
gailo (Minsk) for help in preparation of photographs of the beetles.
Nikolajev G. V., Puntsagdulam Z. Plastinchatousye Mongol'skoi Narodnoi Respubliki // Nasekomye Mon-
golii. – 1984. – 9. – P. 90—294 (in Russian).
|
| id | nasplib_isofts_kiev_ua-123456789-9538 |
| institution | Digital Library of Periodicals of National Academy of Sciences of Ukraine |
| issn | 0084-5604 |
| language | English |
| last_indexed | 2025-11-27T18:22:58Z |
| publishDate | 2001 |
| publisher | Інститут зоології ім. І.І. Шмальгаузена НАН України |
| record_format | dspace |
| spelling | Frolov, A.V. 2010-07-02T10:45:54Z 2010-07-02T10:45:54Z 2001 A Review of the Subgenus Chilothorax of the Genus Aphodius (Coleoptera, Scarabaeidae) Occurring in Mongolia, with Description of New Species / A.V. Frolov // Вестн. зоологии. — 2001. — Т. 35, № 3. — С. 39-45. — Бібліогр.: 1 назв. — англ. 0084-5604 https://nasplib.isofts.kiev.ua/handle/123456789/9538 595.764.1(517.3) The Mongolian species of Aphodius Illiger belonging to the subgenus Chilothorax Motschulsky are reviewed. A. kerzhneri Nikolajev and A. mongolaltaicus Nikolajev are placed in the subgenus Chilothorax. Two new synonyms (A. altaicus Nikolajev = A. puntsagdulamae Nikolajev, syn. n.; A. grafi Reitter = A. boreomongolicus Nikolajev, syn. n.) are established. Three new species are described. A key to Mongolian species of Chilothorax is presented. Дан обзор обитающих в Монголии видов подрода Chilothorax Motschulsky рода Aphodius Illiger. В этот подрод помещены Л. kerzhneri Nikolajev и Л. mongolaltaicus Nikolajev. Установлена синонимия двух видовых названий (Л. altaicus Nikolajev = A. puntsagdulamae Nikolajev, syn. n.; A. grafi Reitter = A. boreomongolicus Nikolajev, syn. n.). Описаны 3 новых вида. Приведена таблица для определения монгольских видов подрода Chilothorax. I would like to thank Dr. Hans Fery (Berlin), Dr. Oleg Kabakov (St. Petersburg), Dr. Izyaslav Kerzhner (St. Petersburg), Dr. David Kral (Praha), Dr. Georgy Nikolajev (Almaty), Dr. Karla Schneider (Halle), and Dr. Alexey Solodovnikov (St. Petersburg) for valuable help. Special thanks are due to Dr. Paul Lago (University of Mississippi) for linguistic review of the manuscript and to Mr. Kirill Dowgailo (Minsk) for help in preparation of photographs of the beetles. en Інститут зоології ім. І.І. Шмальгаузена НАН України Фауна и систематика A Review of the Subgenus Chilothorax of the Genus Aphodius (Coleoptera, Scarabaeidae) Occurring in Mongolia, with Description of New Species Обзор представителей подрода Chilothorax рода Aphodius (Coleoptera, Scarabaeidae) Монголии с описанием новых видов Article published earlier |
| spellingShingle | A Review of the Subgenus Chilothorax of the Genus Aphodius (Coleoptera, Scarabaeidae) Occurring in Mongolia, with Description of New Species Frolov, A.V. Фауна и систематика |
| title | A Review of the Subgenus Chilothorax of the Genus Aphodius (Coleoptera, Scarabaeidae) Occurring in Mongolia, with Description of New Species |
| title_alt | Обзор представителей подрода Chilothorax рода Aphodius (Coleoptera, Scarabaeidae) Монголии с описанием новых видов |
| title_full | A Review of the Subgenus Chilothorax of the Genus Aphodius (Coleoptera, Scarabaeidae) Occurring in Mongolia, with Description of New Species |
| title_fullStr | A Review of the Subgenus Chilothorax of the Genus Aphodius (Coleoptera, Scarabaeidae) Occurring in Mongolia, with Description of New Species |
| title_full_unstemmed | A Review of the Subgenus Chilothorax of the Genus Aphodius (Coleoptera, Scarabaeidae) Occurring in Mongolia, with Description of New Species |
| title_short | A Review of the Subgenus Chilothorax of the Genus Aphodius (Coleoptera, Scarabaeidae) Occurring in Mongolia, with Description of New Species |
| title_sort | review of the subgenus chilothorax of the genus aphodius (coleoptera, scarabaeidae) occurring in mongolia, with description of new species |
| topic | Фауна и систематика |
| topic_facet | Фауна и систематика |
| url | https://nasplib.isofts.kiev.ua/handle/123456789/9538 |
| work_keys_str_mv | AT frolovav areviewofthesubgenuschilothoraxofthegenusaphodiuscoleopterascarabaeidaeoccurringinmongoliawithdescriptionofnewspecies AT frolovav obzorpredstaviteleipodrodachilothoraxrodaaphodiuscoleopterascarabaeidaemongoliisopisaniemnovyhvidov AT frolovav reviewofthesubgenuschilothoraxofthegenusaphodiuscoleopterascarabaeidaeoccurringinmongoliawithdescriptionofnewspecies |