A New Species of Hahnia (Araneae, Hahniidae) from Bulgaria and Crimean Peninsula close to H. nava
Hahnia kovblyuki sp. n. is described from forest habitats in Western Bulgaria and compared to its closest species, H. nava (Blackwall, 1841), using morphology and simple morphometry. Previous records of H. nava from Crimea are formally re-assigned to H. kovblyuki sp. n., and issues in the taxonomy o...
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| author | Indzhov, S. |
| author_facet | Indzhov, S. |
| author_institution_txt_mv | [
{
"author": "S. Indzhov",
"institution": "Faculty of Biology, Sofi a University “St. Kliment Ohridski”"
}
] |
| author_sort | Indzhov, S. |
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| collection | OJS |
| datestamp_date | 2026-06-29T16:20:36Z |
| description | Hahnia kovblyuki sp. n. is described from forest habitats in Western Bulgaria and compared to its closest species, H. nava (Blackwall, 1841), using morphology and simple morphometry. Previous records of H. nava from Crimea are formally re-assigned to H. kovblyuki sp. n., and issues in the taxonomy of the Hahnia nava complex are briefly discussed. |
| doi_str_mv | 10.15407/zoo2026.03.215 |
| first_indexed | 2026-06-30T01:00:36Z |
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DOI 10.15407/zoo2026.03.215
UDC 595.44(497.2)
A NEW SPECIES OF HAHNIA (ARANEAE,
HAHNIIDAE) FROM BULGARIA AND CRIMEAN
PENINSULA CLOSE TO H. NAVA
S. Indzhov
Faculty of Biology, Sofia University “St. Kliment Ohridski”,
8 Dragan Tsankov Blvd, 1164
E-mail: saindzhov@gmail.com
S. Indzhov (https://orcid.org/0000-0002-8937-9373)
urn:lsid:zoobank.org:pub:5E537FA0-F999-4413-8A6A-1F1F005B550E
A New Species of Hahnia (Araneae, Hahniidae) from Bulgaria and Crimean Peninsula close
to H. nava. Indzhov, S. — Hahnia kovblyuki sp. n. is described from forest habitats in West-
ern Bulgaria and compared to its closest species, H. nava (Blackwall, 1841), using morphology
and simple morphometry. Previous records of H. nava from Crimea are formally re-assigned to
H. kovblyuki sp. n., and issues in the taxonomy of the Hahnia nava complex are briefly discussed.
Ke y words : Crimea, Balkan Peninsula, species complex, morphometry, forest habitats, spiders,
taxonomy.
Introduction
The genus Hahnia C. L. Koch, 1841 contains 13 described European species accord-
ing to the World Spider Catalog (2025). These are small spiders which inhabit the
ground layer of various habitats, exhibiting some specialization (species profiles at
Harm, 1966, Kovblyuk et al., 2017). The males have relatively similar copulatory or-
gans, unlike the mostly intricate and specific epigynal structures of the females (Figs
at Harm, 1966, Kovblyuk et al., 2017). However, species complexes with sibling spe-
cies are known from Europe, such as Hahnia ononidum Simon, 1875 and Hahnia
thymorum Ledoux, 2014 (Ledoux, 2014). Hahnia nava (Blackwall, 1841) is a wide-
spread species with a Transpalaearctic distribution (WSC 2025), reaching eastwards
to Japan (Ichikawa, 2024). It is accordingly found in Bulgaria, where it mostly inhab-
its open, grassland habitats (see material listed below), which coincides well with the
Fauna and Systematics Zoodiversity, 60(3): 215–224, 2026
© Publisher Publishing House "Akademperiodyka" of the NAS of Ukraine, 2026. The article is
published under an open access license CC BY-NC-ND (https://creativecommons.org/licenses/
by-nc-nd/4.0/)
ISSN 2707-725X. Zoodiversity. 2026. Vol. 60, No. 3
S. Indzhov
ISSN 2707-725X. Zoodiversity. 2026. Vol. 60, No. 3
216
information given by Harm (1966). However, Kovblyuk et al. (2017), in their treat-
ment of Crimean Hahnia, reported H. nava predominantly from forest habitats, in-
cluding the leaf litter of Fagus forests. In March 2023, a single male Hahnia was
found in Western Bulgaria, in Lozenska Mts., under a stone in a beech forest. It could
not be identified with certainty using the available European literature, but its simi-
larity to Hahnia nava did not remain unnoticed. A direct comparison with unambig-
uous males from the latter species showed a marked difference in the size of the
pedipalps. The following two years, more material was collected at the same spot,
with two more males being found under stones, while the majority of the individuals
were found while sifting and turning older layers of beech litter. The consistent dif-
ferences in the morphology of both sexes and in habitat preferences showed that this
is a separate species, which is described herein.
Material and Methods
Individual spiders were collected while turning stones, searching through vegetation
near ground level, or examining leaf litter, and preserved in approximately 75% etha
nol. Epigynes and palps were separated using syringe needles, and the epigynes were
cleared at room temperature in 80% lactic acid for 2–5 days. The materials were ex-
amined under a Bresser Advance ICD stereomicroscope and a Levenhuk 50L Plus
monocular microscope. Photos of genitalia were taken using a Samsung Galaxy A54
attached to the monocular microscope’s eyepiece and stacked using Picolay (Cypion-
ka, 2025) and subsequently edited using GIMP (2025). Photos of the habitus were
taken through the eyepiece of the Bresser stereomicroscope.
Measurements were made using the diameter of the eyefield as reference (2.5 mm
at 64 and 1 mm at 160 times magnification) and calculated from the pixels using the
program GIMP (2025). All measurements are in mm. Scatter plot was made using R
(2022) in R Studio (2026).
The holotype, two male paratypes and four female paratypes are deposited at
the deposited at the National Museum of Natural History, Sofia (NMNHS). Ad-
ditional paratypes will be deposited in the author’s collection until they are also
transferred to other institution collections in the future; some non-type speci-
mens are in Institute of Biodiversity and Ecosystem Research at the Bulgarian
Academy of Sciences (IBER).
Terminology follows Harm (1966) and Zhang et al. (2011).
Abbreviations: CF, cymbial furrow; Em, embolus; MA, median apophysis; Mts.,
mountain(s).
Results
Family Hahniidae Bertkau, 1878
Genus Hahnia C. L. Koch, 1841
Type species: Hahnia pusilla C. L. Koch, 1841, by subsequent designation of Thorell
(1869).
A New Species of Hahnia (Araneae, Hahniidae) from Bulgaria and Crimean Peninsula
ISSN 2707-725X. Zoodiversity. 2026. Vol. 60, No. 3
217
Hahnia kovblyuki sp. n.
urn:lsid:zoobank.org:act:1409700E-683E-4B67-8AA8-C97476E9319C
Hahnia nava (nec Blackwall, 1841): Kovblyuk et al., 2017: 313, Figs 4–5 [misidentification].
Mater i a l . Type. Bulgaria: Holotype {: Lozen Mts., above Lozen Monastery [42.584° N, 23.521°
E], Fagus forest, underneath a log, 07.03.2023 (S. Indzhov) (NMNHS). Paratypes: 1 {, same data as
holotype, underneath a stone, 04.03.2024; 2 {, 3 }, same data as holotype, sifting beech leaf litter and
under stones, 31.01.2025; 2 {, 4 }, same data as holotype, sifting beech leaf litter, 28.02.2025 (S. In-
dzhov) (NMNHS). Non-type: West Balkan Mts., Gara Lakatnik [43.085° N, 23.378° E], Carpinus for-
est, underneath a stone, 17.10.2025, 1 } (S. Indzhov); West Rhodopes Mts., Chokmanovo [41.547° N,
24.735° E], Fagus-Pinus-Picea forest, 15.11.2025, 1 { (V. Vassilev) (IBER).
Diagnosis . The species is morphologically similar to Hahnia nava (Blackwall,
1841) but can be separated by the following traits: the male cymbium length is ap-
proximately 0.42–0.46 mm (approximately 0.35–0.38 mm in H. nava); the cymbium
length:width ratio is below 1.47 (above 1.50 in H. nava); the cymbial furrow nearly
reaches the tip of the cymbium and ends at a right angle (it reaches below the middle
of the median apophysis and has a rounded ending in H. nava); the embolus origi-
nates at 2 o’clock and runs far from the tegulum in its retrobasal quarter (it originates
at 3 o’clock and runs closer to the tegulum in H. nava); the female copulatory ducts
are longer and usually have larger posterior loops than in the closest species (shorter
ducts and smaller loops in H. nava); the entrance portion of the copulatory ducts is
more strongly pigmented ; the primary spermathecae are approximately 0.065–
0.070 mm in diameter (approximately 0.045 mm in H. nava); the preferred habitat is
humid leaf litter in Fagus forests (open, often dry grasslands for H. nava); the overall
colouration is somewhat paler than in H. nava, especially the prosoma and legs.
Fig. 1. Hahnia nava, male palp, ventral view — a; Hahnia kovblyuki sp. n., male palp, ventral
view — b; Hahnia kovblyuki sp. n., male palp, retrolateral view — c. Scale bar 0.5. Abbreviations:
CF, cymbial furrow; Em, embolus; MA, median apophysis. The distance between the asterisks
denotes the measured cymbium length and the distance between the x symbols denotes the mea-
sured width (measured at the end of the membranous part of Em transversely, see Diagnosis)
a b c
S. Indzhov
ISSN 2707-725X. Zoodiversity. 2026. Vol. 60, No. 3
218
Description
B oth sexes , overa l l appearance, spines
Carapace light brown, with unclear radial striations (Fig. 6). Legs greyish in fresh
specimens, with lighter joints, turning light brown in stored specimens. Opist-
hosoma grey, sometimes with unclear chevrons, ventral side somewhat paler.
Sexual dimorphism is minimal; males are slightly darker, with a shorter head
region and proportionally somewhat longer front legs (for the latter, see meas-
urements). Leg spines are non-dimorphic, as follows (see Table 1).
Male
Measurements (N=3).Total length 1.51–1.68. Carapace length 0.67–0.72. Leg
measurements (male holotype with carapace length 0.71). Femur I 0.52, Patel-
Fig. 2. Hahnia kovblyuki sp. n., dissected epigyne, ventral view — a; Hahnia kovblyuki sp. n., vul-
va, dorsal view — b; Hahnia nava, dissected epigyne, ventral view — c; Hahnia nava, vulva, dorsal
viewb — d. Scale bar 0.2. Abbreviations: CO, copulatory opening; PL, posterior loop; Sp, spermathecae
a
c
b
d
Table 1. Spinulation of Hahnia kovblyuki sp. n.
Segment\Leg I II III IV
Femur 1d1p 1d 1d 1d
Patella 1d 1d 1d 1d
Tibia 1d 1d1v 1d1r2v 1d1p1r2v
Metatarsus none none 1p1r1v 1p1r1v
Tarsus none none 1v none
Abbre v i at ions : d — dorsal; p — prolateral; r — retrolateral; v — ventral.
A New Species of Hahnia (Araneae, Hahniidae) from Bulgaria and Crimean Peninsula
ISSN 2707-725X. Zoodiversity. 2026. Vol. 60, No. 3
219
la I 0.21, Tibia I 0.41, Metatarsus I 0.36, Tarsus I 0.32, Femur II 0.49, Patella II 0.21,
Tibia II 0.40, Metatarsus II 0.32, Tarsus II 0.31, Femur III 0.45, Patella III 0.19, Tibia
III 0.34, Metatarsus III 0.30, Tarsus III 0.31, Femur IV 0.57, Patella IV 0.21, Tibia IV
0.46, Metatarsus IV 0.47, Tarsus IV 0.35.
Palp. Cymbium ovoid, with a tapering tip, somewhat asymmetrical. Cymbial
furrow nearly reaching the tip of the cymbium, ending at a right angle (Figs 1, b;
3, b). Tegulum flat, lacking setose areas, its retrodistal corner irregularly cut off. Em-
bolus filiform, originating at 2 o’clock, with a membranous velum-like area near the
base. Median apophysis originating in the middle of the cut-off section of the tegu-
lum, lamellose, somewhat curved, dilated apically, with a truncated tip. Tibial apo-
physis is long and thin, curved backwards. Patellar apophysis is thin, nearly straight,
but with a hook-shaped tip, and bulged at the base.
Female
Measurements (N=3).Total length 1.92–2.09. Carapace length 0.73–
0.80. Leg measurements (female paratype with carapace length 0.71). Femur I
0.53, Patella I 0.26, Tibia I 0.41, Metatarsus I 0.34, Tarsus I 0.29, Femur II 0.47,
Patella II 0.23, Tibia II 0.35, Metatarsus II 0.25, Tarsus II 0.25, Femur III 0.55,
Fig. 3. Top row, Hahnia kovblyuki sp. n. male pedipalp. Bottom row, Hahnia nava, male pedipalp,
retrolateral view — a, c; ventral view — b, d. Scale bar 0.5
0.5 mm
a
c
b
d
S. Indzhov
ISSN 2707-725X. Zoodiversity. 2026. Vol. 60, No. 3
220
Patella III 0.24, Tibia III 0.32, Metatarsus III 0.34, Tarsus III 0.29, Femur IV 0.65,
Patella IV 0.25, Tibia IV 0.45, Metatarsus IV 0.42, Tarsus IV 0.34.
Epig yne and vulva. Epigyne flat, with no external structures other than two
obliquely situated copulatory openings forming a V (Fig. 2, a). Copulatory ducts are
long, thin, and form several loops (Figs 2, 4, 5), and the entrance portion is more heavily
pigmented. Primary spermathecae spherical to slightly ovoid, rather big. Glandular
structures are visible anteriorly of the dilated entrance parts of the copulatory ducts.
Phenolog y. Adult specimens from both sexes were collected on snowless days in
autumn and winter, from October until the first half of March. It is possible, however,
that their maturity period is even longer, as no targeted searches were conducted in other
parts of the year. Kovblyuk et al. (2017) also show activity in summer — June, August and
September, which perhaps indicates a nearly year-round adult presence.
General distr ibution. Bulgaria, Ukraine (Crimean Peninsula).
Etymolog y. Named after the Ukrainian arachnologist Mykola Kovblyuk
(V. I. Vernadsky Taurida National University, Simferopol, Crimea, Ukraine), who
first illustrated this species.
Hahnia nava (Blackwall, 1841)
Hahnia nava: Harm, 1966; Almquist, 2006 (see WSC (2025) for a complete list of references).
Mater i a l . Non-type. Bulgaria. Central Balkan Mts., above Zlatitsa, Govedarnika
(42.7470° N, 24.1570° E), dry montane pasture, underneath a stone,13.06.2025, 1 } (S. In-
dzhov) (Collection S. Indzhov); Chepan Mts., Chepan ecoalley (42.9480° N, 22.9435° E),
xerothermic slope, underneath stones, 27.03.2021, 2 } (S. Indzhov) (Collection S. Indzhov);
Chepan Mts., road towards Dragoman swamp (42.9373°N, 22.9483° E), roadside, pasture,
underneath a stone, 25.04.2021, 1 {, 1 } (S. Indzhov) (Collection S. Indzhov); Chepan Mts.,
Chepan ecoalley (42.9480° N, 22.9435° E), xerothermic slope, underneath stones, 14.03.2023,
Fig. 4. Top row, Hahnia kovblyuki sp., n., vulva. Bottom row, Hahnia nava: vulva, ventral view —
a, c, dorsal view — b, d. Scale bar 0.2
0.2 mm
a
c
b
d
A New Species of Hahnia (Araneae, Hahniidae) from Bulgaria and Crimean Peninsula
ISSN 2707-725X. Zoodiversity. 2026. Vol. 60, No. 3
221
Fig. 5. Four epigynes of Hahnia kovblyuki sp. n. in situ. Scale bar 0.2
Fig. 6. Four epigynes of Hahnia nava in situ: a — Zlatitsa; b, d — Chepan Mts.; c — Dolni Rak-
ovets. Scale bar 0.2
0.2 mm
2 {, 1 } (S. Indzhov) (Collection S. Indzhov); Chepan Mts., Chepan ecoalley (42.9477° N,
22.9372° E), dry grassland-Quercus grove margin, underneath a stone, 09.03.2025, 1 { (S.
Indzhov) (Collection S. Indzhov); Radomir Basin, Dolni Rakovets (42.4720° N, 23.0223° E),
dry grassland, in grass, 13.05.2025, 1 } (S. Indzhov) (Collection S. Indzhov); Sofia Basin,
near Aldomirovtsi swamp (42.8998° N, 23.0041° E), xerothermic grassland, underneath
stones and in grass tufts, 19.02.2023, 2 {, 1 } (S. Indzhov) (Collection S. Indzhov); Vitosha
a
c
b
d
0.2 mm
a
c
b
d
S. Indzhov
ISSN 2707-725X. Zoodiversity. 2026. Vol. 60, No. 3
222
Mts., Platoto above Kumata Hut (42.5893° N, 23.2527° E), subalpine grassland & Juniperus,
in grass, 16.10.2024, 1 } (S. Indzhov) (Collection S. Indzhov).
Remarks. As this species has been adequately described in the taxonomic literature
(Harm, 1966; Almquist, 2006, etc.), no new description is presented. For comparative
remarks, see the diagnosis of Hahnia kovblyuki sp. n. as well as the key below.
Key to Hahnia nava and Hahnia kovblyuki sp. n. (Hahnia hauseri Brignoli, 1978 excluded)
1 Males............................................................................................................................................... 3
— Females........................................................................................................................................... 4
2 Cymbium length > 0.40 (0.42–0.46), male cymbium length : width ratio < 1.47. Legs
and carapace light greyish brown. Forest species................................... H. kovblyuki sp. n.
— Сymbium length < 0.40 (0.35–0.38), cymbium length : width ratio > 1.50 and above. Legs
and carapace usually strongly pigmented, dark brown. Open habitats......................H. nava
3 Diameter of spermathecae > 0.055 (0.065–0.070), posterior loops often large. Carapace
and legs light greyish brown. Forest species................................................ H. kovblyuki sp. n.
— Diameter of spermathecae < 0.055 (≈0.045), posterior loops small but variable.
Legs and carapace usually strongly pigmented, dark brown. Inhabitant of open habi-
tats. ...............................................................................................................................H. nava
Fig. 7. Hahnia kovblyuki sp. n., habitus. Left, female, right, male. Scale bar 1 mm
1 mm
Fig. 8. Scatter plot showing mea-
surements of male cymbium in
Hahnia nava (N = 5) and Hahnia
kovblyuki sp. n. (N = 4) Squares —
Hahnia nava, circles — Hahnia
kovblyuki sp. n. Filled figures indi-
cate material examined in the cur-
rent paper, empty square is Hahnia
nava from Harm (1966) and emp-
ty circle is Hahnia kovblyuki sp. n.
from Kovblyuk et al. (2017)
C
ym
bi
um
w
id
th
, m
m
Cymbium length, mm
A New Species of Hahnia (Araneae, Hahniidae) from Bulgaria and Crimean Peninsula
ISSN 2707-725X. Zoodiversity. 2026. Vol. 60, No. 3
223
Discussion
Although frequently cited in taxonomic literature (WSC 2025), Hahnia nava appears
to belong to a species complex that includes less-known taxa such as H. kovblyuki sp.
n. and Hahnia hauseri Brignoli, 1978. The latter species is currently known only from
females (Brignoli, 1978); however, it is likely widespread in the Western Mediterrane-
an, as Barrientos (1985) and Lecigne et al. (2025) recorded morphologically similar
females with short posterior loops. A definitive interpretation of these populations re-
mains challenging without additional data on males beyond the descriptions provided
by Barrientos (1985), which is beyond the scope of the present study.
The identity of the male specimen illustrated by Harm (1966) as H. pusilla
C. L. Koch, 1841 remains unclear. The curved median apophysis (MA) and the
embolus originating at approximately 2 o’clock suggest that it belongs to the
Hahnia nava complex; in contrast, H. pusilla possesses a straight MA and an
embolus originating at approximately 5 o’clock (Almquist, 2006). This specimen
is unlikely to be H. kovblyuki sp. n. because its cymbium length is only approxi-
mately 0.35 mm (corrected from 3.5 mm) and it lacks the prominent cymbial
furrow typical of the new species, appearing more similar to H. nava in several
aspects. However, its length:width ratio of approximately 1.48 is more consistent
with H. kovblyuki sp. n. Whether this record represents a misidentified H. nava,
a small specimen of H. kovblyuki sp. n., or an undescribed member of this com-
plex from Central European forests remains unknown, as the original illustra-
tion does not permit a more definitive assessment.
Variability in pedipalp size is evident in both H. kovblyuki sp. n. and H. nava,
with the cymbial length of the latter ranging from 0.35 to 0.38 mm. While literature
data (Harm, 1966; Almquist, 2006) report values around 0.37 mm, our measure-
ments of Bulgarian material (N = 4) show that three males possess a cymbium length
of approximately 0.35 mm, and only one specimen from Dragoman reaches 0.38 mm.
Since this size variation is present in both species and occurs within single localities,
it is interpreted as intraspecific variation.
This variability implies that structural characters — specifically the cymbium
length:width ratio and the extent of the cymbial furrow — are diagnostically more
reliable than absolute measurements alone. Although no overlap in absolute palp
size has been observed in the studied material, the ratio remains a stable and robust
character for distinguishing these sibling species.
In conclusion, the discovery of H. kovblyuki sp. n. highlights the taxonomic com-
plexity within the Hahnia nava species complex in the Western Palaearctic. Given the
subtle morphological differences and potential for overlapping absolute measurements,
accurate identification must rely on a combination of genital proportions and habitat
data. Further research involving molecular analysis and broader geographical sam-
pling across Central Europe and the Mediterranean will be essential to fully clarify the
boundaries of this complex and the status of related populations.
Acknowledgements. The author is thankful to Vassil Vassilev (Sofia, Bulgaria)
for providing an additional specimen and location of Hahnia kovblyuki sp. n. and
also to Graeme Lyons (Brighton, United Kingdom) and Frederik Fischer (Køben-
havn, Denmark) for discussions on the habitat preferences of Hahnia nava. Addi-
S. Indzhov
ISSN 2707-725X. Zoodiversity. 2026. Vol. 60, No. 3
224
tional thanks go to Valeriy Korneyev (I. I. Schmalhausen Institute of Zoology, Na-
tional Academy of Sciences of Ukraine, Kyiv, Ukraine) for the comprehensive advice
on the paper structure, as well as to Anna Hirna (Institute of Ecology of the Car-
pathians NAS of Ukraine, Lviv, Ukraine) and an anonymous reviewer for providing
helpful feedback to the first draft of the paper.
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zootaxa.2912.1.1
Received 22 March 20206
Accepted 30 June 2026
|
| id | oai:ojs.akademperiodyka.org.ua:article-936 |
| institution | Zoodiversity |
| keywords_txt_mv | keywords |
| language | English |
| last_indexed | 2026-06-30T01:00:36Z |
| publishDate | 2026 |
| publisher | Publishing House "Akademperiodyka" of the National Academy of Sciences of Ukraine |
| record_format | ojs |
| resource_txt_mv | ojsakademperiodykaorgua/83/5a7de543aefc5575a939cf23dc751c83.pdf |
| spelling | oai:ojs.akademperiodyka.org.ua:article-9362026-06-29T16:20:36Z A New Species of Hahnia (Araneae, Hahniidae) from Bulgaria and Crimean Peninsula close to H. nava Indzhov, S. Crimea Balkan Peninsula species complex morphometry forest habitats spiders taxonomy Hahnia kovblyuki sp. n. is described from forest habitats in Western Bulgaria and compared to its closest species, H. nava (Blackwall, 1841), using morphology and simple morphometry. Previous records of H. nava from Crimea are formally re-assigned to H. kovblyuki sp. n., and issues in the taxonomy of the Hahnia nava complex are briefly discussed. Publishing House "Akademperiodyka" of the National Academy of Sciences of Ukraine 2026-04-27 Article Article application/pdf https://ojs.akademperiodyka.org.ua/index.php/Zoodiversity/article/view/936 10.15407/zoo2026.03.215 Zoodiversity; Vol. 60 No. 3 (2026): Zoodiversity Zoodiversity (Vestnik Zoologii); Том 60 № 3 (2026): Zoodiversity 2707-7268 2707-725X 10.15407/zoo2026.03 en https://ojs.akademperiodyka.org.ua/index.php/Zoodiversity/article/view/936/394 Copyright (c) 2026 Simeon Indzhov |
| spellingShingle | Indzhov, S. A New Species of Hahnia (Araneae, Hahniidae) from Bulgaria and Crimean Peninsula close to H. nava |
| title | A New Species of Hahnia (Araneae, Hahniidae) from Bulgaria and Crimean Peninsula close to H. nava |
| title_full | A New Species of Hahnia (Araneae, Hahniidae) from Bulgaria and Crimean Peninsula close to H. nava |
| title_fullStr | A New Species of Hahnia (Araneae, Hahniidae) from Bulgaria and Crimean Peninsula close to H. nava |
| title_full_unstemmed | A New Species of Hahnia (Araneae, Hahniidae) from Bulgaria and Crimean Peninsula close to H. nava |
| title_short | A New Species of Hahnia (Araneae, Hahniidae) from Bulgaria and Crimean Peninsula close to H. nava |
| title_sort | new species of hahnia (araneae, hahniidae) from bulgaria and crimean peninsula close to h. nava |
| topic_facet | Crimea Balkan Peninsula species complex morphometry forest habitats spiders taxonomy |
| url | https://ojs.akademperiodyka.org.ua/index.php/Zoodiversity/article/view/936 |
| work_keys_str_mv | AT indzhovs anewspeciesofhahniaaraneaehahniidaefrombulgariaandcrimeanpeninsulaclosetohnava AT indzhovs newspeciesofhahniaaraneaehahniidaefrombulgariaandcrimeanpeninsulaclosetohnava |