Підтверджена популяція клокички перистої (Staphylea pinnata) на горі Радич (Зовнішні Східні Карпати, Україна)
In May 2025, after 145 years, we successfully confirmed a population of rare and relic species, European bladdernut (Staphylea pinnata), on Mt. Radych (Outer Eastern Carpathians) near the village Rozheve, Sambir district, Lviv Region (Ukraine). The collected materials are deposited at the LWS herbar...
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M.M. Gryshko National Botanical Garden of the NAS of Ukraine
2025
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Plant Introduction| _version_ | 1860145174874685440 |
|---|---|
| author | Novikov, Andriy Piechnik, Łukasz Sup-Novikova, Mariia |
| author_facet | Novikov, Andriy Piechnik, Łukasz Sup-Novikova, Mariia |
| author_sort | Novikov, Andriy |
| baseUrl_str | https://www.plantintroduction.org/index.php/pi/oai |
| collection | OJS |
| datestamp_date | 2025-12-27T15:10:16Z |
| description | In May 2025, after 145 years, we successfully confirmed a population of rare and relic species, European bladdernut (Staphylea pinnata), on Mt. Radych (Outer Eastern Carpathians) near the village Rozheve, Sambir district, Lviv Region (Ukraine). The collected materials are deposited at the LWS herbarium. |
| doi_str_mv | 10.46341/PI2025013 |
| first_indexed | 2025-09-17T09:33:47Z |
| format | Article |
| fulltext |
Plant Introduction, 107, 21–25 (2025)
© The Authors. This content is provided under CC BY 4.0 license.
SHORT COMMUNICATION
Confirmed population of European bladdernut (Staphylea pinnata) on
Radych Mt. (Outer Eastern Carpathians, Ukraine)
Andriy Novikov 1, Łukasz Piechnik 2, Mariia Sup-Novikova 3
1 State Museum of Natural History, National Academy of Sciences of Ukraine, Teatralna str. 18, 79008 Lviv, Ukraine;
novikoffav@gmail.com
2 W. Szafer Institute of Botany, Polish Academy of Sciences, Lubicz str. 46, 31-512 Kraków, Poland; l.piechnik@botany.pl
3 Ukrainian Catholic University, Stryiska str. 29A, 79026 Lviv, Ukraine
Received: 10.07.2025 | Accepted: 29.07.2025 | Published online: 31.07.2025
Abstract
In May 2025, after 145 years, we successfully confirmed a population of rare and relic species, European
bladdernut (Staphylea pinnata), on the Mt. Radych (Outer Eastern Carpathians) near the village Rozheve,
Sambir district, Lviv Region (Ukraine). The collected materials are deposited at the LWS herbarium.
Keywords: Staphylea pinnata, distribution, Ukraine, Radych Mt.
https://doi.org/10.46341/PI2025013
UDC 581.95
Authors’ contributions: AN – validation, investigation, resources, writing – original draft. LP – supervision, conceptualization,
methodology, validation, investigation, resources, writing – original draft. MSP – resources, visualization, writing – original draft.
Funding: No special financial funds were involved in realizing this study.
Competing Interests: The authors declare no conflict of interest.
European bladdernut, Staphylea pinnata L.,
(Staphyleaceae Martinov) is a relic deciduous
shrub naturally distributed from southern,
south-eastern and central Europe, Caucasus
and Anatolian Peninsula including Türkiye,
Georgia, Armenia, and Azerbaijan (Piechnik
et al., 2024; POWO, 2025). There is also an
uncertain report of S. pinnata for North Iran
and Syria (Melnyk, 1995; Melnyk et al., 2009).
In Ukraine, S. pinnata is a rare plant
occurring on the northeastern limit of its
distribution range (Melnyk et al., 2009). It is
listed in the last edition of the Red Book of
Ukraine (Melnyk et al., 2009) and, besides
some concerns, including the stability of its
population and lack of threatment in most
of the distribution range (Didukh, 2016), it
has been approved for a new edition in 2021
(MEPNRU, 2021). Here it sporadically appears
in the western and central parts of the country,
where it participates in the broadleaved
forests and subforestal communities (Melnyk,
1995; Derevenko, 2004; Shynder, 2018).
The report of S. pinnata from steppe zone
of Ukraine, i.e., Donetsk Region (Kotov, 1955;
Meusel et al., 1978), appeared to be related to
cultivated plants (Shynder, 2018). Recently,
S. pinnata was reported for Hurivka Zakaznyk
in Kirovohrad Region (Trotner, 2024a, 2024b),
but this is an artificial forest in the steppe
zone and the european bladdernut was most
probably introduced there.
ISSN 1605-6574, e-ISSN 2663-290X
https://creativecommons.org/licenses/by/4.0/
https://orcid.org/0000-0002-0112-5070
https://orcid.org/0000-0002-3958-7393
https://orcid.org/0000-0002-8542-3605
22 Plant Introduction • 107
Novikov et al.
Melnyk et al. (2009) mapped S. pinnata
for the Zakarpattia Region (e.g., Yulivska
Hora Mt. near the village Diula, Kholmovets
zakaznyk near the the village Kholmovets,
and the outskirts of Mukachevo). However,
only locality from Yulivska Hora Mt. has
been recently confirmed (iNaturalist, 2023).
Specimens collected from Yulivska Hora Mt.
are also deposited in the Herbarium of the
State Museum of Natural History of the NAS of
Ukraine (LWS 063174 and LWS 063175 collected
by I.V. Vaynagiy in 1957). Unfortunately, we
could not survey other mapped localities of
S. pinnata in the Zakarpattia Region due to
war-related travel restrictions.
Hence, one of the westernmost confirmed
natural localities of S. pinnata from Ukraine
was Mt. Khomets in Lviv (Znesinnya Regional
Landscape Park). Staphylea pinnata has
been collected on the northern slopes of
Mt. Khomets in 2024 by us. Respective
specimens were deposited at the LWS
herbarium (vouchers 119101, 119103, and
119106). There is a herbarium of S. pinnata
collected toward the west from Lviv, from
Dobromyl outskirts (LWS 063171, LWS 063172,
and LWS 063173, all collected by M.F. Boyko in
1967). However, we were not able to localize
the Dobromyl locality as it was referenced
to Medytska Mt., which is not mapped and
is not associated with any currently known
toponym. Instead, in May 2025, we successfully
confirmed and collected material from the old
locality in Radych Mt., reported briefly over
a century ago by Kotula (1880). This habitat
remained little known and is usually not cited
in works on the distribution of S. pinnata in
Ukraine.Collected specimens are currently
deposited at the LWS herbarium (vouchers
119107, 119113, and 119115).
The Mt. Radych (519 m a.s.l.) is located
near the Ukrainian-Polish border (close to
Dobromyl) to the north-east from the village
Rozheve, Sambir district, Lviv Region (Fig. 1).
It belongs to one of the last isolated short
island mountainous ridges of the Outer
Eastern Carpathians (two summits only) in the
Stryvihor-Dniester interfluve area (Łanczont
et al., 2019). Considering phytogeographical
regionalization, Mt. Radych belongs to the
Peremyshel-Dobromyl region, the Ciscarpathia
district of the Eastern Carpathians sector
(Tasenkevich, 2004). Considering the physico-
geographical mesoregional division, this
Figure 1. The rough outlines of the Staphylea pinnata population on Mt. Radych.
Plant Introduction • 107 23
Confirmed population of European bladdernut (Staphylea pinnata) on Radych Mt.
Figure 2. Staphylea pinnata on Mt. Radych. A – General view of the habitat; B – young plants are seen
growing directly on the path; C – inflorescence. Photo credists: A. Novikov.
A
B C
24 Plant Introduction • 107
Novikov et al.
mountain belongs to the Przemyśl Foothills
(Solon et al., 2018) or the Peremyshel-
Dobromyl Highland (Novikov, 2021).
The top of the mountain ridge is covered
with dense beech-hornbeam-oak forest with a
strong presence of hazelnut. Staphylea pinnata
occurs there in undergrowth at the top of
Mt. Radych and also goes down to northeastern
slopes at least to 400 m elevation, covering ca.
68 ha of area (49.57307, 22.85092 – 49.56583,
22.87663; Fig. 1). On the southwestern slopes
of the ridge, the species was not observed.
It is the most densely represented along
the old pathway at the ridge’s top and under
open gaps in the forest canopy. The nature of
the habitat is typical for S. pinnata, which in
Ukraine demonstrate the highest population
density on forest edges and along forest roads
(Melnyk, 1995; Shynder, 2018). All ontogenetic
stages, including seedlings, are present. Most
of the plants do not exceed 2–2.5 m in height;
however, some plants reaching 3–4 m were
observed (Fig. 2).
Hence, a little-known habitat of S. pinnata,
which is one of the westernmost in Ukraine
was confirmed. This discovery significantly
supplements the knowledge about the range
of the species and encourages us to continue
searching for S. pinnata in the area of Western
Ukraine.
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Plant Introduction • 107 25
Confirmed population of European bladdernut (Staphylea pinnata) on Radych Mt.
Підтверджена популяція клокички перистої (Staphylea pinnata) на горі Радич
(Зовнішні Східні Карпати, Україна)
Андрій Новіков 1, Лукаш Пєхнік 2, Марія Суп-Новікова 3
1 Державний природознавчий музей, Національна академія наук України, вул. Театральна, 18, Львів,
79008, Україна; * novikoffav@gmail.com
2 Інститут ботаніки імені В. Шафера, Польська академія наук, вул. Любіч, 46, Краків, 31-512, Польща;
l.piechnik@botany.pl
3 Український католицький університет, вул. Стрийська, 29A, Львів, 79026, Україна
У травні 2025 року, через 145 років, нам вдалося успішно підтвердити присутність популяції
рідкісного та реліктового виду рослин – клокички перистої (Staphylea pinnata) на горі Радич (Зовнішні
Східні Карпати) поблизу села Рожеве Самбірського району Львівської області (Україна). Зібрані
матеріали зберігаються в гербарії LWS.
Ключові слова: Staphylea pinnata, поширення, Україна, гора Радич
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| spelling | oai:ojs2.plantintroduction.org:article-16632025-12-27T15:10:16Z Confirmed population of European bladdernut (Staphylea pinnata) on Radych Mt. (Outer Eastern Carpathians, Ukraine) Підтверджена популяція клокички перистої (Staphylea pinnata) на горі Радич (Зовнішні Східні Карпати, Україна) Novikov, Andriy Piechnik, Łukasz Sup-Novikova, Mariia In May 2025, after 145 years, we successfully confirmed a population of rare and relic species, European bladdernut (Staphylea pinnata), on Mt. Radych (Outer Eastern Carpathians) near the village Rozheve, Sambir district, Lviv Region (Ukraine). The collected materials are deposited at the LWS herbarium. У травні 2025 року, через 145 років, нам вдалося успішно підтвердити присутність популяції рідкісного та реліктового виду рослин – клокички перистої (Staphylea pinnata) на горі Радич (Зовнішні Східні Карпати) поблизу села Рожеве Самбірського району Львівської області (Україна). Зібрані матеріали зберігаються в гербарії LWS. M.M. Gryshko National Botanical Garden of the NAS of Ukraine 2025-07-31 Article Article application/pdf https://www.plantintroduction.org/index.php/pi/article/view/1663 10.46341/PI2025013 Plant Introduction; No 107 (2025); 21-25 Інтродукція Рослин; № 107 (2025); 21-25 2663-290X 1605-6574 en https://www.plantintroduction.org/index.php/pi/article/view/1663/1571 Copyright (c) 2025 Andriy Novikov, Łukasz Piechnik, Mariia Sup-Novikova http://creativecommons.org/licenses/by/4.0 |
| spellingShingle | Novikov, Andriy Piechnik, Łukasz Sup-Novikova, Mariia Підтверджена популяція клокички перистої (Staphylea pinnata) на горі Радич (Зовнішні Східні Карпати, Україна) |
| title | Підтверджена популяція клокички перистої (Staphylea pinnata) на горі Радич (Зовнішні Східні Карпати, Україна) |
| title_alt | Confirmed population of European bladdernut (Staphylea pinnata) on Radych Mt. (Outer Eastern Carpathians, Ukraine) |
| title_full | Підтверджена популяція клокички перистої (Staphylea pinnata) на горі Радич (Зовнішні Східні Карпати, Україна) |
| title_fullStr | Підтверджена популяція клокички перистої (Staphylea pinnata) на горі Радич (Зовнішні Східні Карпати, Україна) |
| title_full_unstemmed | Підтверджена популяція клокички перистої (Staphylea pinnata) на горі Радич (Зовнішні Східні Карпати, Україна) |
| title_short | Підтверджена популяція клокички перистої (Staphylea pinnata) на горі Радич (Зовнішні Східні Карпати, Україна) |
| title_sort | підтверджена популяція клокички перистої (staphylea pinnata) на горі радич (зовнішні східні карпати, україна) |
| url | https://www.plantintroduction.org/index.php/pi/article/view/1663 |
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